Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Aquatic Coleoptera (Beetles) of Michigan

Introduction

The information contained herein is still preliminary and should not be considered authoratative.

Certain groups of aquatic beetles have been well surveyed in our state, and several systematic monographs have appeared for several important families (e.g., Dytiscidae) that occur in our state. Still, the enormous number of Coleopteran species that may occur in our state means that to fashion completed species lists for all the families listed to the left will require additional work. More information regarding aquatic beetles and Coleoptera in genera can be found at the Coleoptera Home Page.

Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

Adephaga Schellenberg, 1806

    DYTISCIDAE Leach, 1815 (Predaceous Diving Beetles)

    Photo of Bidessonotus sp. adult from Honduras, Copyright © 2006 Ethan Bright
    Bidessontus sp. from Honduras

    The Nearctic region has been relatively well surveyed, and most of this information has been recently included in a new publication by Larson et al. (2000). This list and nomenclature is largely based on that work as well as provisional records in the UMMZ and the Smithsonian (NMNH web page database).

      Acilius Leach, 1817 (Subfamily Dytiscinae; Tribe Aciliini)
        Acilius athabascae Larson, 1975 - MN, boreal, may occur in Isle Royale NP, nw UP (Larson et al. 2000: 813-814)
        Acilius fraternus (Harris, 1828) Dytiscus (orig.) - Hilsenhoff 1975: 273; see also Larson et al. 2000: 812
        Acilius mediatus (Say, 1823) Dytiscus (orig.) - Hilsenhoff 1975: 273; see also Larson et al. 2000: 807-808
        Acilius semisulcatus Aubé, 1838 - Hilsenhoff 1975: 273-274; Larson et al. 2000: 808-810
        Acilius sylvanus Hilsenhoff, 1975 - Hilsenhoff 1975: 271-273; Larson et al. 2000: 811-812

      Agabetes Crotch, 1873 (Subfamily Agabetinae)
        Agabetes acuductus (Harris, 1828) - IN, ON, WI, expect in LP (Hilsenhoff 1992: 64-65; Larson et al. 2000: 56-58)
          (syn.) Colyembetes acauductus Harris, 1828
          (syn.) Agabus regulosis Aubé, 1838

      Agabus Leach, 1817 (Subfamily Colymbetinae, Tribe Agabini)
        Agabus aeruginosus Aubé, 1838 - Larson et al. 2000: 596-597
        Agabus ambiguus (Say, 1823) Colymbetes (orig.) - Larson 1989: 879-883; Larson et al. 2000: 625-626
        Agabus antennatus Leech, 1939 - MN, ON, WI, widespread north (Hilsenhoff 1993c: 181; Larson et al. 2000: 580-582)
          (syn.) Agabus clavatus LeConte
        Agabus anthracinus Mannerheim, 1852 - Larson 1989: 896-898; Larson et al. 2000: 635-637
        Agabus bicolor (Kirby, 1837) Colymbetes (orig.) - Larson 1991: 1269-1271, 1314; Larson et al. 2000: 679-681
        Agabus bifarius (Kirby, 1837) Colymbetes (orig.) - Larson et al. 2000: 589-591
        Agabus canadensis Fall, 1922 - MN, ON, WI, expect in UP (Larson 1991: 1264-1266, 1313; Hilsenhoff 1993c: 183; Larson et al. 2000: 675-676)
        Agabus confinis (Gyllenhal, 1808) Dytiscus (orig.) - Larson 1991: 1287-1289, 1311; Larson et al. 2000: 692-694
        Agabus confusus (Blatchley, 1910) Rhantus (orig.) - WI (MI? found in south central WI: Hilsenhoff 1993c: 183)
          (syn.) Agabus ampulus Fall, 1922
        Agabus discolor (Harris, 1828) Colymbetes (orig.) - Larson 1991: 1272-1275, 1315; Larson et al. 2000: 683-684
        Agabus disintegratus (Crotch, 1873) Gaurodytes (orig.) - UMMZ (see also Hilsenhoff 1993c: 184; Larson et al. 2000: 600-602)
        Agabus erichsoni Gemminger & Harold, 1868 - Larson et al. 2000: 506-508
        Agabus falli (Zimmermann, 1934) - IN, MN, ON, WI (Hilsenhoff 1993c: 185; Larson et al. 2000: 616-617)
          (syn.) Agabus semipunctatus Sharp, 1882
          (syn.) Agabus sharpi Fall, 1922
          (comb.) Gaurodytes falli Zimmermann, 1934
        Agabus flavovittatus Larson and Wolfe, 1998 - IL, IN, OH, ON (expect in s. MI; see Larson et al. 2000: 542)
        Agabus fuscipennis (Paykull, 1798) Dytiscus (orig.) - ON (boreal, may occur in UP) (Hilsenhoff 1993c: 185; Larson 1994: 155-156, 180; Larson et al. 2000: 6583-654)
        Agabus gagates Aubé, 1838 - record at NMNH (web page database); see also Larson et al. 2000: 508-509
        Agabus immaturus Larson, 1991 - Larson 1991: 1263-1264, 1313; Larson et al. 2000: 673-675
        Agabus inscriptus (Crotch, 1873) Gaurodytes (orig.) - Larson 1991: 1281-1284, 1316; Larson et al. 2000: 688-689
        Agabus larsoni Fery and Nilsson, 1993 - ON (Larson et al. 2000: 509-510)
        Agabus leptapsis (LeConte, 1878) Gaurodytes (orig.) - NMNH (web page database); see also Larson 1989: 892-894; Larson et al. 2000: 632-634
        Agabus obtusatus (Say, 1823) Colymbetes (orig.) - IN, OH, ON (Hilsenhoff 1993c: 187; Larson et al. 2000: 585-586)
        Agabus opacus Aubé, 1837 - UMMZ, NMNH (web page database - as A. pseudoconfertus Wallis); Larson 1996: 631-633, 664; Larson et al. 2000: 521-523
          (syn.) Agabus sharpi Fall, 1922
          (syn.) Gaurodytes mimmi Shalberg, 1875
          (syn.) Agabus sibericus Sharp, 1882
          (syn.) Agabus gelidus Fall, 1926
          (syn.) Agabus pseudoconfertus Wallis, 1926
          (syn.) Agabus sachalinensis Kamiya, 1938
        Agabus phaeopterus (Kirby, 1837) Colymbetes (orig.) - Larson 1991: 1260-1262, 1312; Larson et al. 2000: 671-673
        Agabus punctatus Melsheimer, 1844 - Larson et al. 2000: 594-595
        Agabus punctulatus Aubé, 1838 - IL, IN, MN, ON, WI (Hilsenhoff 1993c: 188; Larson et al. 2000: 609-610)
        Agabus semipunctatus (Kirby, 1837) Colymbetes (orig.) - Larson et al. 2000: 605-606
        Agabus semivittatus LeConte, 1852 - UMMZ
        Agabus seriatus (Say, 1823) Colymbetes (orig.) - Larson et al. 2000: 559-561
        Agabus subfuscatus Sharp, 1882 - Larson 1991: 1271-1272, 1317; Larson et al. 2000: 681-683
        Agabus thomsoni (Sahlberg, 1871) Gaurodytes (orig.) - Larson 1991: 1255-1257, 1309; Larson et al. 2000: 666-668
        Agabus wasastjernae (Sahlberg, 1824) Dytiscus (orig.) - Larson 1996: 629-631, 664; Larson et al. 2000: 519-521

      Bidessonotus Régimbart, 1895 (Subfamily Hydroporinae, Tribe Bidessini)
        Bidessonotus inconspicuus (LeConte, 1855) Hydroporus (orig.) - UMMZ; see also Larson et al. 2000: 105-106
        Bidessonotus pulicarius (Aubé, 1838) Hydroporus (orig.) - IN, ON (Larson et al. 2000: 107

      Celina Aubé, 1837 (Subfamily Hydroporinae, Tribe Methlini)
        Celina hubbelli Young, 1979 - Larson et al. 2000: 90-91

      Colymbetes Clairville, 1806 (Subfamily Colymbetinae, Tribe Colymbetini)
        Colymbetes exaratus LeConte, 1862 - IL, MN, WI, may occur in the UP (Zimmerman 1981: 13-16 (as C. e. exaratus), Larson et al. 2000: 770)
          (syn.) Colymbetes sculptilis exaratus, Young & Severin, 1956
          (syn.) Colymbetes sculptilis (in part), Larson, 1975
          (syn.) Colymbetes rugipennis Sharp, 1882
        Colymbetes paykulli Erichson, 1837 - Zimmerman 1981: 40-41; Larson et al. 2000: 765-766
          (syn.) Colymbetes longulus LeConte, 1862
        Colymbetes sculptilis Harris, 1829 - Zimmerman 1981: 11, 18-20, 22-27; Larson et al. 2000: 767-769

      Copelatus Erichson, 1832 (Subfamily Copelatinae)
        Copelatus chevrolati Aubé, 1838 - Larson et al. 2000: 48, 50-51
          (syn.) Copelatus renovatus Guignot, 1952
        Copelatus glyphicus (Say, 1823) - Larson et al. 2000: 51-52

      Coptotomus Say, 1834 (Subfamily Colymbetinae, Tribe Coptotomini)
        Coptotomus longulus LeConte, 1852 - Hilsenhoff 1980: 465-466 (as C. lenticus); Larson et al. 2000: 734-736
          (syn.) Coptotomus lenticus Hilsenhoff, 1980
        Coptotomus loticus Hilsenhoff, 1980 - IL, IN, ON, WI (Hilsenhoff 1980: 467-468; Larson et al. 2000: 736-737)

      Cybister Curtis, 1827 (Subfamily Dytiscinae, Tribe Cybistrini)
        Cybister fimbriolatus (Say, 1823) Dytiscus (orig.) - UMMZ, NMNH (web page database); see also Larson et al. 2000: 832-833

      Desmopachria Babington, 1841 (Subfamily Hydroporinae, Tribe Hyphydrini)
        Desmopachria convexa (Aubé, 1838) Hydroporus (orig.) - UMMZ, NMNH (web page database); Larson et al. 2000: 99-100; see also Young 1981: 3; Hilsenhoff 1994: 277

      Dytiscus Linnaeus, 1758 (Subfamily Dytiscinae, Tribe Dytiscini)
        Dytiscus alaskanus Balfour-Browne, 1944 - MN, ON (near Sault Ste. Marie), WI, expect in UP and n. LP (Roughley 1990: 481-483, 500; Hilsenhoff 1993a: 41; Larson et al. 2000: 792-793)
          (syn.) Dytiscus parvulus Motschulsky, 1852
        Dytiscus carolinus Aubé, 1838 - Roughley 1990: 446-447, 453
          (syn.) Dytiscus fasciventris; auctorim, nec Say, 1824
          (syn.) ?Dytiscus fasciventris, Folkerts 1978
          (syn.) Dytiscus sp., Young 1954
        Dytiscus circumcinctus Ahrens, 1811 - Roughley 1990: 477-481, 498; Larson et al. 2000: 791-792
          (syn.) Dytiscus flavocinctus Hummel, 1823
          (syn.) Dytiscus angustatus Stephens 1828
          (syn.) Dytiscus dubius Gyllenhal, 1827
          (syn.) Dytiscus ooligbuckii Kirby, 1837
          (syn.) Dytiscus circumscriptus Boisduval & Lacordaire, 1835
          (syn.) Dytiscus anxius Mannerheim, 1843
          (syn.) Dytiscus fusco-striatus Motschulsky, 1853
          (syn.) Dytiscus confusus Motschulsky, 1860
          (misid.) Dytiscus albinicus; auctorum, nec Motschulsky, 1859
          (midid.) Dytiscus dauricus; auctorum, nec Gebler, 1832
        Dytiscus cordieri Aubé, 1838 - Roughley 1990: 456-458, 463; Larson et al. 2000: 789-790
        Dytiscus dauricus Gebler, 1832 - Roughley 1990: 483-486, 501; Larson et al. 2000: 793-795
        Dytiscus fasciventris Say, 1824 - Roughley 1990: 447-449, 454; Larson et al. 2000: 786-788
        Dytiscus harrisii Kirby, 1837 - Roughley 1990: 433-434, 440; Larson et al. 2000: 782-783
        Dytiscus hybridus Aubé, 1838 - Roughley 1990: 434-435, 441; Larson et al. 2000: 783-785
        Dytiscus verticalis Say, 1823 - Roughley 1990: 431-433, 439; Larson et al. 2000: 780-782

      Graphoderus Dejean, 1833 (Subfamily Dytiscinae, Tribe Aciliini)
        Graphoderus fasciocollis (Harris, 1828) Dytiscus (orig.) - Larson et al. 2000: 821-822
        Graphoderus liberus (Say, 1825) Dytiscus (orig.) - Larson et al. 2000: 817-818
        Graphoderus manitobensis Wallis, 1933 - IL, WI, expect in western UP (Hilsenhoff 1993a: 47-48; Larson et al. 2000: 822-823)
        Graphoderus occidentalis Horn, 1883 - NMNH (web page database); see also Larson et al. 2000: 820-821
        Graphoderus perplexus Sharp, 1882 - Larson et al. 2000: 818-819

      Heterosternuta Strand, 1935 (Subfamily Hydroporinae, Tribe Hydroporini)
        Heterosternuta pulcher (LeConte, 1855) - OH, ON, MN (Hilsenhoff 1995a: 4; Larson et al. 2000: 235-236)
        Heterosternuta wickhami (Zaitzev, 1907) Hydroporus (orig.) - UMMZ; see also Larson et al. 2000: 232-234

      Hydaticus Leach, 1817 (Subfamily Dytiscinae, Tribe Hydaticini)
        Hydaticus aruspex Clark, 1864 - Roughley and Pengelly 1981: 260-267, 300 (as H. modestus); Larson et al. 2000: 800-801
          (syn.) Hydaticus modestus Sharp, 1882
        Hydaticus piceus LeConte, 1863 - Roughley and Pengelly 1981: 267-268, 302; Larson et al. 2000: 802-803

      Hydrocolus (=Hydroporus oblitus-group Fall, 1923) Roughley & Larson 2000 (Subfamily Hydroporinae, Tribe Hydroporini)
        Hydrocolus paugus (Fall, 1923), Hydroporus (orig.) - Larson et al. 2000: 414-416
        Hydrocolus persimilis (Crotch, 1873), Hydroporus (orig.) Larson et al. 2000: 416-417
          (syn.) Hydroporus aequus Fall, 1923
          (syn.) Hydroporus acadianus Balfour-Brown, 1948
        Hydrocolus rybyae (Larson, 1975) Hydroporus (orig.) - ON, WI (boreal, expect in UP; Hilsenhoff 1995a: 16; Larson et al. 2000: 422-423)
        Hydrocolus stagnalis (Gemminger & Harold, 1868) Hydroporus (orig.) - Larson et al. 2000: 426-428
          (syn.) Hydroporus colaris LeConte, 1855

      Hydroporus Clairville, 1806 (Subfamily Hydroporinae, Tribe Hydroporini)
        Hydroporus americanus Aubé, 1838 - Larson et al. 2000: 293-294
        Hydroporus appalachius Sherman, 1913 - MN, ON, expect in UP (Larson et al. 2000: 353-355)
          (syn.) Hydroporus signatus Sharp, 1882
          (syn.) Hydroporus occidentalis Gordon and Post, 1965
        Hydroporus badiellus Fall, 1923 - UMMZ, NMNH (web page database); holotype from Marquette (Larson et al. 2000: 301-303)
        Hydroporus brevicornis Fall, 1917 - UMMZ, NMNH (web page database); Larson et al. 2000: 299-300
        Hydroporus columbianus Fall, 1923 - UMMZ, NMNH (web page database); Larson et al. 2000: 328-330
        Hydroporus dentellus Fall, 1917 - UMMZ, NMNH (web page database); Larson et al. 2000: 294-296
        Hydroporus despectus Sharp, 1882 - UMMZ, NMNH (web page database); see also Larson et al. 2000: 397-399
          (syn.) Hydroporus rusticus Sharp, 1882
        Hydroporus dichrous Melsheimer, 1844 - UMMZ, NMNH (web page database); Larson et al. 2000: 289-291
        Hydroporus fuscipennis Schaum, 1868 - record at Albertson College of Idaho Collection; see also Larson et al. 2000: 345-348
          (syn.) Hydroporus puberulus Mannerheim, 1853
          (syn.) Hydroporus criniticoxis Larson, 1975
        Hydroporus gossei Larson and Roughley, 2000 - Larson et al. 2000: 405-407
        Hydroporus larsoni Nilsson, 1984 - MN, ON, WI (boreal, Hilsenhoff 1995a: 10-11; Larson et al. 2000: 350-353)
        Hydroporus melsheimeri Fall, 1917 - Larson et al. 2000: 291-293
        Hydroporus morio Aubé, 1838 - Larson et al. 2000: 340-342
          (syn.) Hydroporus melanocephalus Gyllenhal, 1808
          (syn.) Hydroporus caliginosus LeConte, 1850
        Hydroporus niger Say, 1823 - Larson et al. 2000: 306-308
          (syn.) Hydroporus modestus Aubé, 1838
          (syn.) Hydroporus latifrons Sharp, 1882
        Hydroporus nigellus Mannerheim, 1853 - widespread boreal, expect in UP (Larson et al. 2000: 389-391)
          (syn.) Hydroporus sp. near despectus, Larson, 1975: 312
        Hydroporus notabilis LeConte, 1850 - Larson et al. 2000: 303-306
          (syn.) Hydroporus arcticus Thompson, 1854
          (syn.) Hydroporus niger Anderson, 1962
        Hydroporus obscurus Sturm, 1835 - Larson et al. 2000: 296-299
          (syn.) Hydroporus hockingi Larson, 1975
        Hydroporus puberulus LeConte, 1850 - ON, WI (boreal, Hilsenhoff 1995a: 12-13; Larson et al. 2000: 337-339)
          (syn.) Hydroporus glabriusculus Fall, 1923
          (syn.) Hydroporus levanderi Sahlberg, 1888
        Hydroporus rectus Fall, 1923 - Larson et al. 2000: 403-404
        Hydroporus rufilabris Sharp, 1882 - Larson et al. 2000: 311-312
          (syn.) Hydroporus somnus Fall, 1923
        Hydroporus rufinasus Mannerheim, 1852 - Hilsenhoff 1995a: 13; Larson et al. 2000: 335-337
        Hydrporus signatus Mannerheim, 1853 - Larson et al. 2000: 312-315
          (syn.) Hydroporus inornatus Sharp, 1882
          (syn.) Hydroporus niger Sharp, 1882
        Hydroporus striola Gyllenhal, 1827 - Larson et al. 2000: 365-367
          (syn.) Hydroporus subtonsus LeConte, 1855
        Hydroporus tartaricus LeConte, 1850 - UMMZ; see also Larson et al. 2000: 385-389
          (syn.) Hydroporus elusivus Gordon, 1981
        Hydroporus tenebrosus LeConte, 1850 - Larson et al. 2000: 395-397
          (syn.) Hydroporus luridipennis LeConte, 1850
          (syn.) Hydroporus lutulentus LeConte, 1855
        Hydroporus tristis (Paykull, 1798) Dytiscus - Larson et al. 2000: 343-345
          (syn.) Hydroporus varians LeConte, 1850
          (syn.) Hydroporus ruficapillus Mannerheim, 1852

      Hydrovatus Motschulsky, 1853 (Subfamily Hydroporinae, Tribe Hydrovatini)
        Hydrovatus pustulatus (Melsheimer, 1844) Hygrotus - Larson et al. 2000: 94-95; see Biström 1997 if other taxa in MI, or Young 1956, 1963
          (syn.) Hydrovatus indianensis Blatchley, 1910

      Hygrotus Stephens, 1828 (Subfamily Hydroporinae, Tribe Hydroporini)
        Hygrotus acaroides (LeConte, 1855) Hydroporus (orig.) - NMNH (web page database); see also Larson et al. 2000: 150-152
          (comb.) Coelambus acaroides winnipeg Wallis, 1926
        Hygrotus compar (Fall, 1919) Coelambus (orig.) - Larson et al. 2000: 167-169
        Hygrotus dissimilis Gemminger and Harold, 1868 - Larson et al. 2000: 171-173
          (syn.) Hydroporus dispar LeConte, 1850
        Hygrotus falli (Wallis, 1924) Coelambus (orig.) - Anderson 1983: 190; Hilsenhoff 1994: 282; Larson et al. 2000: 180-182
        Hygrotus farctus (LeConte, 1855) Hydroporus (orig.) - Larson et al. 2000: 152-153
        Hygrotus impressopunctatus (Schaller, 1783) Dytiscus (orig.) - Anderson 1983: 190-191; Larson et al. 2000: 194-196
          (syn.) Hydroporus similis Kirby, 1837
        Hygrotus laccophilinus (LeConte, 1878) Hydroporus (orig.) - UMMZ, record at Albertson College of Idaho Collection; Larson et al. 2000: 157-159
        Hygrotus marklini (Gyllenhal, 1813) Hyphydrus (orig.) - Anderson 1983: 183 (as H. canadensis); Hilsenhoff 1994: 283; Larson et al. 2000: 162-164
          (syn.) Coelambus canadensis Fall, 1919
          (syn.) Coelambus virgo Fall, 1919
        Hygrotus nubilus (LeConte, 1855) Hydroporus (orig.) - Anderson 1983: 189; Larson et al. 2000: 178-180
        Hygrotus patruelis (LeConte, 1855) Hydroporus (orig.) - UMMZ; see also Anderson 1983: 194; Larson et al. 2000: 176-178
          (syn.) Hydroporus discoideus LeConte, 1855
          (syn.) Coelambus borealis Fall, 1937
          (syn.) Coelambus quebecensis Brown, 1930
        Hygrotus picatus (Kirby, 1837) Hydroporus (orig.) - Anderson 1983: 191-192; Larson et al. 2000: 196-198
        Hygrotus punctilineatus (Fall, 1919) - questionable UMMZ (check); see Larson et al. 2000: 186-188
        Hygrotus sayi Balfour-Browne, 1944 - Larson et al. 2000: 154-156
          (syn.) Laccophilus punctatus Say, 1824, type lost
        Hygrotus sellatus (LeConte, 1866) Hydroporus (orig.) - Anderson 1983: 182-183; Hilsenhoff 1994: 285; Larson et al. 2000: 164-166
        Hygrotus suturalis (LeConte, 1850) Hydroporus (orig.) - type specimen from Eagle Harbor, MI; Anderson 1983: 182; Hilsenhoff 1994: 285; see also Larson et al. 2000: 161-162
          (syn.) Hydroporus ovoideus LeConte, 1850
        Hygrotus sylvanus (Fall, 1917) Coelambus (orig.) - UMMZ; see also Larson et al. 2000: 156-157
        Hygrotus turbidus (LeConte, 1855) Hydroporus (orig.) - Larson et al. 2000: 159-161
        Hygrotus unguicularis (Crotch, 1874) Hydroporus (orig.) - MB, QB, widespread boreal, expect in UP (Larson et al. 2000: 192-194)

      Ilybius Erichson, 1832 (Subfamily Colymbetinae, Tribe Agabini)
        Ilybius angustior (Gyllenhal, 1808) Dytiscus (orig.) - Larson 1987: 382-387; Larson et al. 2000: 718-720
        Ilybius biguttulus (Germar, 1824) Dytiscus (orig.) - Larson 1987: 367-371; Larson et al. 2000: 710-712; Merritt et al. 2002: 155
          (syn.) Ilybius laramaeus LeConte, 1859
        Ilybius confusus Aubé, 1838 - Larson 1987: 376-379; Larson et al. 2000: 715-716
          (syn.) Ilybius denikei Wallis 1933
        Ilybius discendens Sharp, 1882 - Larson 1987: 350, 360-362; Larson et al. 2000: 704-706
        Ilybius fraterculus LeConte, 1862 - Larson 1987: 364-367; Hilsenhoff 1993c: 194; Larson et al. 2000: 708-709
        Ilybius ignarus (LeConte, 1862) Colymbetes (orig.) - Larson et al. 2000: 712-713
        Ilybius incarinatus Zimmermann, 1928 - Larson 1987: 363, 373-374, 376; Hilsenhoff 1993c: 195; Larson et al. 2000: 713-715
        Ilybius oblitus Sharp, 1882 - Larson 1987: 379-382; Hilsenhoff 1993c: 195; Larson et al. 2000: 716-717
        Ilybius picipes (Kirby, 1837) Colymbetes (orig.) - NMNH (web page database); Larson 1987: 388-391 (Isle Royale); Larson et al. 2000: 722-723
        Ilybius pleuriticus LeConte, 1850 - Larson 1987: 393-396; Larson et al. 2000: 725-726
          (syn.) Ilybius inversus Sharp, 1882
        Ilybius subaeneus Erichson, 1837 - UMMZ; see also Larson 1987: 396-399; Larson et al. 2000: 726-728
          (syn.) Ilybius virianeus Crotch, 1873
          (syn.) Ilybius suffusus Crotch, 1873

      Laccophilus Leach, 1817 (Subfamily Laccophilinae)
        Laccophilus biguttatus Kirby, 1837 - Larson et al. 2000: 67-68
          (syn.) Laccophilus inconspicuus Fall, 1917
        Laccophilus fasciatus Aubé, 1838 - Hilsenhoff 1992: 67
          (syn.) Laccophilus rufus Melsheimer, 1844
          (syn.) Laccophilus terminalis Sharp, 1882
        Laccophilus maculosus Say, 1823 - Larson et al. 2000: 64-65
          (syn.) Laccophilus decipiens LeConte, 1852
          (syn.) Laccophilus shermani Leech, 1944
        Laccophilus proximus Say, 1823 - Larson et al. 2000: 66-67
        Laccophilus undatus Aubé, 1838 - IL, IN, OH, ON, WI (Hilsenhoff 1992: 67-68; see Zimmerman 1970 if in MI

      Laccornis Gozis, 1914 (Subfamily Hydroporinae, Tribe Laccornini)
        Laccornis conoideus (LeConte, 1850) Hydroporus (orig.) - Wolfe and Roughley 1990: 312-313, 343; Larson et al. 2000: 79-81
          (syn.) Agaporus conoideus Zimmermann, 1919
        Laccornis deltoides (Fall, 1923) Agaporus (orig.) - Wolfe and Roughley 1990: 316-317, 343; Larson et al. 2000: 84-85
        Laccornis latens (Fall, 1937) Agaporus (orig.) - Wolfe and Roughley 1990: 315-316, 342; Larson et al. 2000: 82-84

      Liodessus Guignot, 1939 (Subfamily Hydroporinae, Tribe Bidessini)
        Liodessus affinis (Say, 1823) Hydroporus (orig.) - Miller 1998: 299-301; Larson et al. 2000: 113-114
          (comb.) Bidessus affinis (Say, 1823), Sharp 1882 and authors
        Liodessus cantralli (Young, 1953) Bidessus (orig.) - Larson and Roughley 1990: 240; Hilsenhoff 1994: 288; Miller 1998: 310; Larson et al. 2000: 116-117
        Liodessus flavicollis (LeConte, 1855) Hydroporus (orig.) - Larson et al. 2000: 111-113; see also Miller 1998: 311
        Liodessus fuscatus (Crotch, 1873) Hydroporus (orig.) - Larson and Roughley 1990: 238-239; Larson et al. 2000: 119-120
        Liodessus obscurellus (LeConte, 1852) Hydroporus (orig.) - IN, MN, WI (Miller 1998: 303-308; Larson et al. 2000: 115-116)
          (syn.) Hydroporus macularis LeConte, 1852
          (syn.) Hydroporus erythrostomus Mannerheim, 1852
          (syn.) Hydroporus charlotti Clark, 1862
          (syn.) Hydroporus emilianus Clark, 1862
          (syn.) Bidessus nigrinus Casey, 1884
          (syn.) Bidessus affinis microreticulatus (Hatch, 1828) - see Miller 1998 for taxonomic history)

      Lioporeus Guignot, 1950 (Subfamily Hydroporinae, Tribe Hydroporini)
        Lioporeus triangularis (Fall, 1917) - MI? check collection, and Wolfe 1983; OH, WI (Hilsenhoff 1994: 290; Larson et al. 2000: 240-241)

      Matus Aubé, 1836 (Subfamily Colymbetinae, Tribe Matini)
        Matus bicarinatus (Say, 1823) Colymbetes (orig.) - Larson et al. 2000: 741
        Matus ovatus Leech, 1941 - Larson et al. 2000: 739-741

      Nebrioporus Régimbart, 1906 (Subfamily Hydroporinae, Tribe Hydroporini)
        Nebrioporus rotundatus (LeConte, 1863) Hydroporus (orig.) - Larson et al. 2000: 450-452

      Neoporus Guignot, 1931 (Subfamily Hydroporinae, Tribe Hydroporini)
        Neoporus carolinus (Fall, 1917) Hydroporus (orig.) - IN, ON (Larson et al. 2000: 260-261)
        Neoporus clypealis (Sharp, 1882) Hydroporus (orig.) Larson et al. 2000: 258-259
        Neoporus dimidiatus (Gemminger and Harold, 1868), Hydroporus (orig.) - ON, widespread (Larson et al. 2000: 266-268)
          (syn.) Hydroporus semirufus LeConte, 1850
          (syn.) Hydroporus solitarius Sharp, 1882
        Neoporus mellitus (LeConte, 1855) Hydroporus (orig.) - Hilsenhoff 1995a: 19; Larson et al. 2000: 246-247
        Neoporus spurius (LeConte, 1855) Hydroporus (orig.) - Larson et al. 2000: 255-256
        Neoporus sulcipennis (Fall, 1917) Hydroporus (orig.) - ON (Bruce Co.) Larson et al. 2000: 253-255
        Neoporus superioris (Balfour-Browne), 1944 Hydroporus (orig.) - MN, ON, WI, expect in UP (Hilsenhoff 1995a: 20; Larson et al. 2000: 268-270)
          (syn.) Hydroporus superioris LeConte, 1850
        Neoporus tennetum (Wolfe, 1984) Hydroporus (orig.) - Larson et al. 2000: 248-249
        Neoporus undulatus (Say, 1823) Hydroporus (orig.) - UMMZ; (see also Hilsenhoff 1995a: 20-21; Larson et al. 2000: 261-264)
          (syn.) Hydroporus fasciatus Harris, 1828
          (syn.) Hydroporus pubipennis Aubé, 1838
          (syn.) Hydroporus velutinus Aubé, 1838
          (syn.) Hydroporus punctatissimus Aubé, 1838
          (syn.) Hydroporus consimilis LeConte, 1850
          (syn.) Hydroporus scrutator Sharp, 1882
          (syn.) Hydroporus anticus Sharp, 1882
          (syn.) Hydroporus placatus Fall, 1923
        Neoporus vitiosus (LeConte, 1855) Hydroporus (orig.) - Larson et al. 2000: 251-253
        Neoporus vittatus (LeConte, 1855) Hydroporus (orig.) - IL, IN, ON (western), MN, WI (Hilsenhoff 1995a: 21; Larson et al. 2000: 264-266)

      Neoscutopterus Balfour-Browne, 1943 (Subfamily Colymbetinae, Tribe Colymbetini)
        Neoscutopterus angustus (LeConte, 1850) Agabus (orig.) - UMMZ, NMNH (web page database); Larson et al. 2000: 744-745
        Neoscutopterus hornii (Crotch, 1873) Scutopterus (orig.) - UMMZ, NMNH (web page database); Larson et al. 2000: 745746

      Oreodytes Seidlitz, 1887 (Subfamily Hydroporinae, Tribe Hydroporini)
        Oreodytes laevis (Kirby, 1837) Hydroporus (orig.) - widespread northern, may occur in UP (Larson et al. 2000: 460-461)
        Oreodytes scitulus (LeConte, 1855) Hydroporus (orig.) - NMNH (web page database); see also Hilsenhoff 1994: 290; Larson et al. 2000: 466-468

      Rhantus Dejean, 1833 (Subfamily Colymbetinae, Tribe Colymbetini)
        Rhantus binotatus (Harris, 1828) Colymbetes (orig.) - Zimmerman and Smith 1975: 58-68; Larson et al. 2000: 755-756
          (syn.) Colymbetes assimilis Kirby, 1837
          (syn.) Colymbetes divisus Aubé, 1838
          (syn.) Rhantus flavogriseus Crotch, 1873
          (syn.) Rhantus longipes Sharp, 1882
          (syn.) Rhantus plejibus Sharp, 1882
          (syn.) Rhantus obscurus Sharp, 1882
          (syn.) Rantus aequalis Hatch, 1953
        Rhantus consimilis Motschoulsky, 1859 - Larson et al. 2000: 752-753
          (syn.) Colymbetes tostus LeConte, 1866
          (syn.) Rhantus tostus (LeConte, 1866) of authors
          (syn.) Rhantus discedens Sharpt, 1882
        Rhantus gutticollis (Say, 1834) Colymbetes (orig.) - WI, maybe w. UP (Larson et al. 2000: 756-757)
          (syn.) Colymbetes mexicanus Laport (de Castelnau), 1835
          (syn.) Rhantus mexicanus, Leech 1948
          (syn.) Rhantus flavogriseus, Leech, 1848
          (syn.) Rhantus hubbelli Hatch, 1928
          (syn.) Rhantus dominguensis Gemminger & Harold, 1868
          (syn.) Rhantus hoppingi Wallis, 1933
        Rhantus sericans Sharp, 1882 - MN, ON, WI, expect in UP (Zimmerman and Smith 1975: 74-81 (as R. frontalis); Larson et al. 2000: 754-756)
        Rhantus sinuatus (LeConte, 1862) Colymbetes (orig.) - Zimmerman and Smith 1975: 105-108; Larson et al. 2000: 751-753
        Rhantus suturellus (Harris, 1828) Colymbetes (orig.) - Zimmerman and Smith 1975: 93-100; Larson et al. 2000: 759-760
          (syn.) Rhantus bistriatus (Bergstrasser), of authors
          (syn.) Rhantus zimmermanni Wallis, 1833
        Rhantus wallisi Hatch, 1953 - Larson et al. 2000: 758-759
          (syn.) Rhantus suturellus Wallis (nec Harris), 1933
          (syn.) Rhantus bistriatus Bergstrasser, F. Balfour-Browne, 1950

      Sanfilippodytes Franciscolo, 1979 (Subfamily Hydroporinae, Tribe Hydroporini)
        Sanfilippodytes pseudovilis (Young, 1953) Hydroporus (orig.) - holotype from E.S. George Reserve, Livingston Co.; Young 1953; see also Hilsenhoff 1995a: 21-22

      Stictotarsus Zimmermann, 1919 (Subfamily Hydroporinae, Tribe Hydroporini)
        Stictotarsus griseostriatus (DeGeer, 1774) Dytiscus (orig.) - UMMZ, NMNH (web page database); see also Zimmerman and Smith 1975: 657-661; Larson et al. 2000: 442-444
          (comb.) Hydroporus griseostriatus, Aubé, 1883 and authors
          (syn.) Hydroporus catascopium Say, 1823
          (syn.) Hydroporus parallelus Say, 1823
          (syn.) Hydroporus coloradensis Fall, 1923
          (syn.) Hydroporus mathiasi Hatch, 1933
          (syn.) Deronectes suffusus Sharp, 1882

      Thermonectus Dejean, 1837 (Subfamily Dytiscinae)
        Thermonectus basillaris (Harris, 1829) Dytiscus (orig.) - IL, IN, ON, WI (Hilsenhoff 1993a: 52; Larson et al. 2000: 826-827)
        Thermonectus nigrofasciatus ornaticollis Aubé, 1838 - WI, may occur in s. MI (Hilsenhoff 1993a: 52 (as T. ornaticollis)

      Uvarus Guignot, 1939 (Subfamily Hydroporinae, Tribe Bidessini)
        Uvarus granarius (Aubé, 1838) Hydroporus (orig.) - NMNH (web page database); see also Larson et al. 2000: 129-131
        Uvarus falli (Young), 1940 Bidessus - ON (Larson et al. 2000: 131-132)
        Uvarus lacustrus (Say, 1823) Hydroporus (orig.) - UMMZ, NMNH (web page database); see also Larson et al. 2000: 133-134

    GYRINIDAE Latreille, 1802 (Whirligig Beetles)

    Adult Dineutus sp. (Copyright © 2006 Ethan Bright)

      Dineutus MacLeay, 1825
        Dineutus assimilis Kirby, 1837 - UMMZ, NMNH (web page database)
        Dineutus discolor Aubé, 1838 - NMNH (web page database)
        Dineutus emarginatus Say, 1825 - UMMZ; see also Hilsehoff 1990: 83, 85
        Dineutus hornii Roberts, 1895 - UMMZ, NMNH (web page database)
        Dineutus nigrior Roberts, 1895 - UMMZ

      Gyrinus Geoffroy, 1762
        Gyrinus aeneolus LeConte, 1868 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 59-60, 68
        Gyrinus aeratus Stephens, 1835 - ON, NY (expect in UP, n. LP) (Oygur and Wolfe 1990: 38-39, 72)
          (syn.) Gyrinus aeneus Thomson, 1860
          (syn.) Gyrinus thomsoni Zaitsev, 1907
          (syn.) Gyrinus edwardsi Sharp, 1914
          (syn.) Gyrinus instabilis Fall, 1931
        Gyrinus affinis Aubé, 1838 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 30-33, 52
          (syn.) Gyrinus canadensis Régimbart, 1883
        Gyrinus analis Say, 1825 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 62-63,67
        Gyrinus aquiris LeConte, 1868 - UMMZ, NMNH (web page database); Oygur and Wolfe 1990: 66, 68-70
        Gyrinus bifarius Fall, 1922 - UMMZ; Oygur and Wolfe 1990: 40-42, 70
          (syn.) Gyrinus punctellus Ochs, 1949
        Gyrinus borealis Aubé, 1838 - Hilsenhoff 1990: 88; Oygur and Wolfe 1990: 45-46, 50; UMMZ
          (syn.) Gyrinus corpulentus Régimbart, 1883
        Gyrinus conifinis LeConte, 1868 - UMMZ, record at Albertson College of Idaho Collection; Oygur and Wolfe 1990: 54-55, 70
        Gyrinus dubius Wallis, 1926 - northern, may occur on Isle Royale (Oygur and Wolfe 1990: 65, 73)
        Gyrinus dichrous LeConte, 1868 - Oygur and Wolfe 1990: 61-62, 71; UMMZ
        Gyrinus fraterunus Couper, 1865 - Mackinaw Co. (Merritt et al. 2002: 155)
        Gyrinus gehringi Chamberlain, 1929 - Oygur and Wolfe 1990: 66-67
        Gyrinus gibber LeConte, 1868 - Oygur and Wolfe 1990: 48-49, 51; UMMZ
          (syn.) Gyrinus frosti Fall, 1922
          (syn.) Gyrinus floridensis Ochs, 1929
        Gyrinus impressicollis Kirby, 1837 - Oygur and Wolfe 1990: 36, 72
        Gyrinus latilimbus Fall, 1922 - Oygur and Wolfe 1990: 53-54, 71; UMMZ
        Gyrinus lecontei Fall, 1922 - UMMZ; Oygur and Wolfe 1990: 66-68
        Gyrinus maculiventris LeConte, 1868 - Oygur and Wolfe 1990: 64, 67; UMMZ
        Gyrinus marginellus Fall, 1922 - Oygur and Wolfe 1990: 46, 48, 69; UMMZ
        Gyrinus minutus Fabricius, 1798 - Oygur and Wolfe 1990: 25-27, 30-31 (however, this may be G. cavatus Atton, 1990 - check article in Can Ent. that O & W were unaware of); UMMZ
          (syn.) Gyrinus kirbyi Marsham, 1802
        Gyrinus pectoralis LeConte, 1868 - Oygur and Wolfe 1990: 24, 72; Merritt et al. 2002: 155; UMMZ
        Gyrinus pernitidus LeConte, 1868 - NY, WI (Oygur and Wolfe 1990: 51, 56-57)
        Gyrinus piceolus Blatchely, 1910 - IN (known only from Indiana, but might occur in sw. LP) (Oygur and Wolfe 1990: 49, 51)
        Gyrinus picipes Aubé, 1838 - ON, expect in w. UP, Isle Royale (Oygur and Wolfe 1990: 64-65, 73)
        Gyrinus pugionis Fall, 1922 - Oygur and Wolfe 1990: 46, 50; UMMZ
        Gyrinus sayi Aubé, 1838 - Oygur and Wolfe 1990: 65-66, 73; UMMZ
          (syn.) Gyrinus lugens LeConte, 1868
          (syn.) Gyrinus falli Notman, 1919
        Gyrinus ventralis Kirby, 1837 - Oygur and Wolfe 1990: 49-51; Merritt et al. 2002: 155; UMMZ
          (syn.) Gyrinus hatchi Wallis, 1926
        Gyrinus wallisi Fall, 1922 - MN, ON, expect in UP (Oygur and Wolfe 1990: 36-37, 72)
        Gyrinus woodruffi Fall, 1922 - UMMZ

    HALIPLIDAE Aubé, 1836 (Crawling Water Beetles)

    Photos of adult and larva of Brychius hungerfordi (Copyright © 2006 Ethan Bright)

      Brychius Thompson 1859
        Brychius hungerfordi Spangler, 1954 - Endangered Species (collection in the USA illegal without a U.S. federal permit) Spangler 1954; Strand and Spangler 1994; Legge 1997; UMMZ
      Haliplus Latreille 1802
        Haliplius apostolicus Wallis, 1913 - WI
        Haliplus blanchardi Roberts, 1913 - UMMZ
        Haliplus borealis LeConte, 1850 - NMNH (web page database)
        Haliplus canadensis Wallis, 1933 - UMMZ
        Haliplus connexus Matheson, 1912 - UMMZ
        Haliplus cribrarius LeConte, 1850 - Hickman 1929: 414-418; Downie and Arnett 1996
        Haliplus fasciatus Aubé, 1838 - Downie and Arnett 1996
        Haliplus immaculicollis Harris, 1828 - Hickman 1929: 409-414; UMMZ; Merritt et al. 2002: 155
        Haliplus leopardus Roberts, 1913 - IL, IN, OH, WI
        Haliplus longulus LeConte, 1850 - UMMZ
        Haliplus nitens LeConte, 1850 - ON, WI?
        Haliplus ohioensis - IL, IN, OH
        Haliplus pantherinus Aubé, 1838 - Downie and Arnett 1996
        Haliplus ruficollis DeGeer - UMMZ
        Haliplus subguttatus Crotch, 1873 - UMMZ, record at Albertson College of Idaho Collection
        Haliplus tortilipennis Brigham and Sanderson, 1972 - IL (Cook Co.), WI
        Haliplus triopsis Say, 1823 - Hickman 1929: 418-421; UMMZ
        Haliplus variomaculatus - IL (Vermillion Co.)
      Peltodytes Regimbart 1872
        Peltodytes edentulus (LeConte, 1863) - Hickman 1929: 408-409; UMMZ, NMNH (web page database)
        Peltodytes duodecimpunctatus (Say, 1823) - IL, IN, OH, WI
        Peltodytes lengi Roberts, 1913 - Hickman 1929: 400-407; Downie and Arnett 1996
        Peltodytes muticus (LeConte) - UMMZ
        Peltodytes pedunculatus Blatchley - Downie and Arnett 1996 - NMNH (web page database)
        Peltodytes sexmaculatus Roberts - Hickman 1929: 407-408; Downie and Arnett 1996
        Peltodytes tortulosus Roberts - Downie and Arnett 1996

    NOTERIDAE Thomson, 1860 (Burrowing Water Beetles)

    Photo of Hydrocanthus iricolor (Copyright © 2012 Tom Murray, used with permission)
    Copyright © 2012 Tom Murray
      Hydrocanthus Say, 1823
        Hydrocanthus iricolor Say, 1823 - UMMZ
      Pronoterus Sharp, 1882
        Pronoterus semipunctatus (LeConte, 1878) Suphis (orig.) - Downie and Arnett 1996
      Suphisellus Crotch, 1873
        Suphisellus bicolor Say, 1831 Noterus (orig.) - IL, IN, OH
        Suphisellus puncticollis Crotch, 1873 - Downie and Arnett 1996
Polyphaga

    CHRYSOMELIDAE Latreille, 1802 (Leaf Beetles)

    The great majority of species in this very large beetle family are terrestrial, but a few taxa, which have specialized on phytophagy upon Nymphaeaceae and other aquatic plants, have become associated with, or adapted to, aquatic habitats (Jolivet and Hawkeswood 1995). The ecology of aquatic and semiaquatic chrysomelids has been summarized by Jolivet and Verma (2002). Larvae of all aquatic species (Donaciinae) use a pair of specialized hooks at the end of the abdomen to tap oxygen from air-containing chambers in plant tissue. Mandibles of the larvae are adapted to macerate plant tissue, and the maxilliary galea and lacinia are adapted to use pharyngeal suction of plant fluids. Larvae construct pupal chambers on their host plants, which is connected to the plant's air cavities. As in the aquatic-adapted Curculionidae, semiaquatic taxa utilize cuticular waxes and hairs as a respiratory plastron or to aid in swimming.Species citations are based on Downie and Arnett (1996), and this list must be considered provisional as no other literature has been consulted, or musuem specimens examined.

      Disonycha Chevrolat, 1844 (Subfamily Galerucinae, Tribe Alticini)
        [not all species in genus are semiaquatic, and I have tried to omit terrestrial species; on Amaranthaceae]
        Disonycha admirabilis Blatchley, 1924 - IN, OH (Downie and Arnett 1996: 1370)
        Disonycha alternata (Illiger), 1807 Haltica (orig.) - Downie and Arnett 1996: 1370 (on Salix, willow; perhaps periaquatic)
        Disonycha latifrons laticollis Schaeffer, 1919 - Downie and Arnett 1996: 1370 (found in wetlands on goldenrod; perhaps periaquatic)
        Disonycha pensylvanica (Illiger, 1807) Haltica (orig.) - (on Polygonum spp.)
        Disonycha procera Casey, 1884 - Downie and Arnett 1996: 1370 (on Polygonum spp.)
        Disonycha uniguttata (Say, 1824) Altica (orig.) - Downie and Arnett 1996: 1370

      Donaciella Rietter, 1920 (Subfamily Donaciinae)
        Donaciella pubicollis (Suffrian, 1872) Donacia (orig.) - Downie and Arnett 1996: 1289 (on Phragmites spp., reed-grass, and Nymphaea spp., water lilies)
          (orig.) Donacia pubicollis Suffrain, 1872:21
          (syn.) Donacia pubicollis Crotch (1872:21, not Suffrian, 1872:21)

      Donacia Fabricius, 1775 (Subfamily Donaciinae)
        Donacia (Donaciomima) biimpressa Melsheimer, 1847 - Downie and Arnett 1996:1298 (on Carex spp., Scirpus spp., bulrushes)
          (syn.) Donacia aurichalcea Melsheimer, 1847:159)
          (syn.) Donacia torosa LeConte, 1851:313
        Donacia (Donaciomima) caerulea Olivier, 1795 - Downie and Arnett 1996:1296 (on Sagittaria spp.; also on Acorus calamus sweet flag, and Pontederia cordata, pickerel weed); Pine Creek, 10 mi S Battle Creek, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia aequalis Say, 1824:248
          (syn.) Donacia confusa, Lacordaire 1845:109, not LeConte, 1851
          (syn.) Donacia confusa LeConte 1851:313, not Lacordaire, 1845
        Donacia (Donaciomima) cazieri Marx, 1957 - Munising Tourist Park (Kölsch & Pedersen 2008) (Downie and Arnet 1996:1298, on Carox limosa, mud sedge)
        Donacia (Donacia) cincticornis Newman, 1838 - Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies, Potamogeton spp. pondweed, Nymphoides spp., floating heart)
          (syn.) Donacia lucida Lacordaire, 1845:106
          (syn.) Donacia rufipennis Lacordaire, 1845:110
          (syn.) Donacia pulchella LeConte, 1851:312
          (syn.) Donacia antillarum Suffrain, 1864:282
          (syn.) Donacia cincticornis var. tryphera Schaeffer, 1925:81
          (syn.) Donacia cincticornis var. tenuis Schaeffer, 1925:82
        Donacia (Donaciomima) confluenta Say, 1827 - Downie and Arnett 1996:1297 (on Sparganium spp., bur-reeds); Pine Creek, 10 mi S Battle Creek, MI; ditch along Mitchell Lake Rd., MI (Kölsch & Pedersen 2008)
        Donacia (Donaciomima) cuprea Kirby, 1837 - Downie and Arnett 1996:1296 (on Scirpus occidentalis)
          (syn.) Donacia curticollis Knab, 1905:122
        Donacia (Donaciomima) distincta Leconte, 1851 - Downie and Arnett 1996:1298 (on Carex spp. sedges, Scirpus spp., bulrushes)
        Donacia (Donaciomima) fulgens Leconte, 1851- Downie and Arnett 1996:1296-1297 (on Sparganium angustifolium, narrow leafed bur-reed, Pontederia cordata, pickerel weed); small lake near Lake City, MI (Kölsch & Pedersen 2008)
        Donacia (Donaciomima) hirticollis Kirby, 1837 - Downie and Arnett 1996:1296 (on Potamogeton spp., other aquatic plants); small lake near Lake City, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia rudicollis Lacordaire, 1845:108
        Donacia (Donaciomima) liebecki Schaeffer, 1919 - Downie and Arnett 1996:1297 (on Nymphaea spp., Nymphar spp., Orontium aquaticum, floating arum)
          (syn.) Donacia pallipes Lacordaire, 1845:149
        Donacia (Donaciomima) limonia Schaeffer, 1925 - IN, ON, may occur in s. LP (Downie and Arnett 1996:1298; on Carex spp., sedges, and Scirpus spp., bulrushes)
        Donacia (Donaciomima) magnifica Leconte, 1851 - ON, MN, widespread (Downie and Arnett 1996:1297 (on Potamogeton spp., pond weeds, Sagittaria spp., arrow leaf)
        Donacia (Donaciomima) megacornis Blatchley, 1910 - Downie and Arnett 1996:1297 (on Nymphaea spp., Nymphar spp.)
          (syn.) Donacia megatocera Weise 1913:118
        Donacia (Donacia) palmata Olivier, 1795 - Downie and Arnett 1996: 1294 (on Nymphaea spp. and Nuphar spp., water lilies); ; small lake near Lake City, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia claudicans Germar, 1821:173
          (syn.) Donacia palmata angustipes Marx, 1957:237
        Donacia (Donacia) parvidens Schaeffer, 1919 - Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies)
        Donacia (Donacia) piscatrix Lacordaire, 1845 - Downie and Arnett 1996: 1294-1295 (on Nymphaea spp. and Nuphar spp., water lilies); shallow lake E Tacoma Lake, MI (Kölsch & Pedersen 2008)
          (syn.) Donacia carolina Lacordaire, 1845:114
          (syn.) Donacia cuprea Melsheimer, 1847:158, not Kirby, 1837
          (syn.) Donacia alutacea LeConte, 1851:311
          (syn.) Donacia congener LeConte, 1851:310
        Donacia (Donaciomima) porosicollis Lacordaire, 1845 - MN, ON, expect in Isle Royale, w UP (Downie and Arnett 1996: 1298; on Carex spp. sedges, and Scirpus spp. nut rushes)
        Donacia (Donacia) proxima Kirby, 1837 - Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies)
          (syn.) Donacia espiscopalis Lacordaire, 1845:105
          (syn.) Donacia california LeConte, 1861:357
          (syn.) Donacia caerulea, sensu Schaeffer 1925 and Marx 1957
        Donacia (Donaciomima) pubescens Leconte, 1868 - Downie and Arnett 1996:1296 (on Scirpus occidentalis)
        Donacia (Donacia) rufescens Lacordaire, 1845 - IN, OH, ON (Downie and Arnett 1996: 1295 (on Nymphaea spp. and Nuphar spp., water lilies), widespread east)
        Donacia (Donaciomima) rugosa Leconte, 1851 - Downie and Arnett 1996:1298 (on Pontederia cordata, pickerel weed)
          (syn.) Donacia aenea Ahrens, 1810:21 (not Hoppe 1875:44, not Gistel 1857:524)
          (syn.) Donacia quadricollis Say, 1827:282
          (syn.) Donacia aerea Lacordaire, 1845:148 (not Schrank, 1798)
          (syn.) Donacia subtilis magistrigata Mead, 1938:113
        Donacia (Donaciomima) subtilis Kunze, 1818 - Downie and Arnett 1996:1297 (on Sparganium spp., bur-reeds); Kalamazoo River, 4 mi E Douglas, MI; Fawn River, 1 mi E Constantine, MI (Kölsch & Pedersen 2008)
        Donacia (Donacia) texana Crotch, 1873 -Downie and Arnett 1996: 1295-1296 (on Nymphaea spp. water lilies)
          (syn.) Donacia texana var. minor Schaeffer, 1925:72
        Donacia (Donaciomima) tuberculifrons Schaeffer, 1919 - Downie and Arnett 1996:1296 (on Scirpus paludosus, Sparganium spp., Carex spp., Acorus calamus sweet flag)

      Hydrothassa Thomson, 1866
        [Ranunculus and marsh vegetation]
        Hydrothassa boreella Schaeffer, 1928 - widespread, expect northern Michigan (Downie and Arnett 1996: 1346)
        Hydrothassa obliquata (Leconte, 1866) Prasocuris (orig.) - IN, OH (Downie and Arnett 1996: 1346)
        Hydrothassa ovalis (Blatchley, 1910) Prasocuris (orig.) - Downie and Arnett 1996: 1346, on Carex spp.
        Hydrothassa vittata (Olivier, 1808) Helodes (orig.) - IN, OH, widespread (Downie and Arnett 1996: 1346)
          (syn.) Hydrothassa trivittata (Say)
          (syn.) Hydrothassa varipes (LeConte)

      Neohaemonia Székessy, 1941
        Neohaemonia flagellata Askevold, 1988 - IL, IN, ON, MN (Downie and Arnett 1996: 1289; widespread, on Potamogeton spp.)
        Neohaemonia melsheimeri (Lacordaire, 1845) Haemonia (orig.) - Downie and Arnett 1996: 1289; on Potamogeton spp.
        Neohaemonia minnesotensis Askevold, 1988 - Downie and Arnett 1996: 1289; on Potamogeton spp.
        Neohaemonia nigricornis (Kirby, 1837) Haemonia (orig.) - MN, ON, WI, widespread (Downie and Arnett 1996: 1288-1289; on Potamogeton spp.)
          (orig.) Haemonia nigricornis Kirby, 1837:222)
          (syn.) Haemonia americana Guérin-Méneville, 1844:249

      Plateumaris Thomson, 1859
        Plateumaris aurifera (LeConte, 1851) Donacia - MN (Askevold 1991: 155; expect in northern UP, on Carex, Scirpus and Eleocharis)
        Plateumaris diversa (Schaeffer, 1925) Donacia - Askevold 1991: 40, 151 (ESGR, Livinston Co.; on sedges in moist meadows)
          (syn.) Donacia harrisi Blatchley, 1910:1105 (misid.)
        Plateumaris flavipes (Kirby, 1837) - Donacia - Askevold 1991: 156 (on Carex, Scirpus and Eleocharis)
          (syn.) Donacia wallisi Schaeffer, 1925
        Plateumaris frosti (Schaeffer, 1925) - Askevold 1991: 164 (on Carex esp. stricta Lam., Scirpus and Eleocharis), Kölsch & Pedersen 2008: 938 (Soldier Lake [46.349N, -84.867W], Chippewa Co.)
        Plateumaris fulvipes (Lacordaire, 1845) - Askevold 1991: 158 (on Carex, Scirpus and Eleocharis)
          (syn.) Donacia fulvipes Dejean, 1837 (nom.nudum)
        Plateumaris germari (Mannerheim, 1843) - Donacia - Askevold 1991: 157 (on Carex, Scirpus, Juncus and Eleocharis)
          (syn.) Donacia flavipennis Mannerheim, 1843
          (syn.) Donacia dives LeConte, 1851
          (syn.) Donacia serricauda Schaeffer, 1919
          (syn.) Donacia femoralis LeConte, 1851 (misid.)
        Plateumaris metallica (Ahrens, 1810) - Donacia - Askevold 1991: 160 (on Carex, Scirpus, and Juncus affinis R. Brown, adventitious on other plants)
          (syn.) Donacia femoralis, Kirby (1837: 223), Lacordaire, 1845
          (syn.) Donacia cataractae Newman (1838: 391)
          (syn.) Donacia parva Lacordaire (1845: 190)
          (syn.) Donacia indica Melsheimer (1847: 158)
          (syn.) Donacia nana Melsheimer (1847: 160)
          (syn.) Donacia gentilis LeConte (1851: 314)
        Plateumaris nitida (Germar, 1811) - Donacia - Askevold 1991: 163 (on Carex, Scirpus and Eleocharis, adventitious on other plants), Kölsch & Pedersen 2008: 938 (M-115 at Middle Branch River, [44.088N, -85.231W], Osceola Co.)
          (syn.) Donacia emarginata Kirby, 1837, var. pacifica Schaeffer, 1925
          (syn.) Donacia juncina Couper, 1864
        Plateumaris pusilla (Say, 1827) - Donacia - Askevold 1991: 153 (on Carex, Scirpus, Juncus and Eleocharis, adventitious on other plants)
          (syn.) Donacia rugifrons Newman, 1838 (nom.nudum), Lacordaire, 1845
          (syn.) Donacia pyritosa LeConte, 1857
          (syn.) Donacia cuprea Crotch, 1873:31 (misid.)
        Plateumaris robusta (Schaffer, 1919) - Donacia - Askevold 1991: 154 (on Carex, Scirpus and Eleocharis)
        Plateumaris rufa (Say, 1827) - Donacia - Askevold 1991: 152 (on Carex and Scirpus)
          (syn.) Donacia sulcicollis Dejean, 1837 (nom.nudum), Lacordaire, 1845
          (syn.) Donacia affis Kirby, 1837
          (syn.) Donacia chalcea Lacordaire, 1845
          (syn.) Donacia kirbyi Lacordaire, 1845
          (syn.) Donacia jacunda LaConte, 1850 (nom.nudum), 1851
        Plateumaris shoemakeri (Schaeffer, 1925) - Donacia - Askevold 1991: 159 (on Acorus calamus L., accidental or adventitious on other plants)
          (syn.) Donacia flavipes var. shoemakeri Schaeffer, 1925
          (syn.) Donacia flavipes var. lodingi Schaeffer, 1925
          (misid.) Plateumaris chalcea, Askevold, 1990
        Plateumaris schaefferi Askevold, 1991 - Askevold 1991: 162

      Prasocuris Latreille, 1802
        Prasocuris phellandrii (Linnaeus, 1758) Chrysomela (orig.) - IN, OH, ON, widespread (Downie and Arnett 1996: 1345, on Caltha spp. (marsh marigold) and Sium suave (water parsnip)

      Pyrrhalta Joannis, 1866
        [not aquatic]
        Pyrrhalta (Tricholochomaea) alni (Fall, 1924) Galerucella (orig.) - Downie and Arnett 1996:1352 (on Alnis, alder; periaquatic)
        Pyrrhalta (Tricholochomaea) decora (Say, 1824) Galleruca (orig.) - Downie and Arnett 1996 (on Salix, willow; perhaps periaquatic)
        Pyrrhalta (Neogalerucalla) quebecensis (Brown, 1938) Galerucella (orig.) - Downie and Arnett 1996:1350 (on Potentialla palustris, cinquefoil; periaquatic)
        Pyrrhalta (Tricholochomaea) ribicola (Brown, 1938) Galerucella (orig.) - Downie and Arnett 1996:1352 (on Ribes, gooseberry; periaquatic, especially bogs)
        Pyrrhalta (Tricholochomaea) tuberculata (Say, 1824) Galleruca (orig.) - Downie and Arnett 1996:1352, (on Salix, willow; perhaps periaquatic)

    CURCULIONIDAE Latreille, 1802 (True Weevils)

      Photo of Auleutes nebulosus by Tom Murray (Copyright © 2011, used with permission).
      Copyright © 2011 Graham Montgomery, used with permission

      This list is based on Downie and Arnett (1996) and must be considered provisional. No museum specimens or other literature citations have been consulted.

        Auleutes Dietz, 1896, 1802 (Subfamily Ceutorhynchinae)
          (not all species listed herein are aquatic/semiaquatic)
          Auleutes asper (LeConte, 1876) Coeliodes (orig.) - IN
          Auleutes atar (LeConte, 1876) Coeliodes (orig.) - IN
          Auleutes epilobii (Paykull, 1800) Curculio (orig.) - Downie and Arnett 1996
          Auleutes marionis Fall, 1913 - IN
          Auleutes nebulosus (LeConte, 1876) Coeliodes - Downie and Arnett 1996
          Auleutes subfasciatus Dietz, 1896 - Downie and Arnett 1996
          Auleutes tenuipes (LeConte), 1876 Coeliodes - IN, ON

        Bagous Germar, 1817 (Subfamily Bagoinae)
          Bagous americanus LeConte, 1876 - Tanner 1943:19-20 (also B. longirostrus? 18-19); Downie and Arnett 1996:1535
          Bagous atratus Blatchley, 1916 - Tanner 1943:23-24; Downie and Arnett 1996:1535
          Bagous bituberosa LeConte, 1876 - IN (widespread, Tanner 1943:29-30; Downie and Arnett 1996:1536)
          Bagous blanchardi Blatchley, 1916 - Downie and Arnett 1996:1535
          Bagous calvifrons LeConte, 1876 - Tanner 1943:21-22; Downie and Arnett 1996:1535
          Bagous magister LeConte, 1876 - Tanner 1943:22; Downie and Arnett 1996:1535
          Bagous mamillatus Say, 1831 - Tanner 1943:16-18; Downie and Arnett 1996:1534-1535
          Bagous nebulosus LeConte, 1876 - Tanner 1943:25; Downie and Arnett 1996:1535
          Bagous obliquus LeConte, 1876 - Downie and Arnett 1996:1534
          Bagous planatus LeConte, 1876 - Tanner 1943:12; Downie and Arnett 1996:1534
          Bagous puritanus Blatchley, 1916 - IN, IL (Downie and Arnett 1996:1535-1536)
          Bagous pusillus LeConte, 1876 - IL, IN (Downie and Arnett 1996:1536)
          Bagous restrictus LeConte, 1876 - IL, IN, MN, OH, WI (Tanner 1943:28; Downie and Arnett 1996:1535)
          Bagous tanneri O'Brien, 1979 - Downie and Arnett 1996:1535
          Bagous transversus LeConte, 1876 - Tanner 1943:31-32; Downie and Arnett 1996:1535
            (syn.) Bagous carinatus Blatchley, 1925

        Brachybamus Germar, 1836 (Subfamily Erirhininae)
          Brachybamus electus Germar, 1836 - IN (widespread, on Sagittaria; Downie and Arnett 1996:1538)

        Euhrychiopsis Dietz, 1896 (Subfamily Ceutorhynchinae)
          Euhrychiopsis lecontei (Dietz, 1896) Phytobius (orig.) - Creed 1998

        Lissorhopterus LeConte 1876 (Subfamily Erirhininae)
          Lissorhopterus buchanani Kuschel, 1952 - Downie and Arnett 1996:1537
          Lissorhopterus lacustris Kuschel, 1952 - Downie and Arnett 1996:1537
          Lissorhopterus oryzophilus Kuschel, 1952 - Downie and Arnett 1996:1537 (rice water weevil)

        Listronotus Jekel, 1865 (Subfamily Cylydrorhininae)
          [some species listed herein may not be aquatic or semi-/peri-aquatic]
          Listronotus appendiculatus (Boheman, 1842) Listroderes (orig.) - Downie and Arnett 1996:1503
          Listronotus caudatus (Say, 1824) Rhynchaenus (orig.) - Downie and Arnett 1996:1501-1502
          Listronotus delumbis (Gyllenhal, 1834) Listroderes (orig.) - Downie and Arnett 1996:1505
          Listronotus dorsalis (Dietz, 1889) Macrops (orig.) - IN, IL, MN (Downie and Arnett 1996:1504)
          Listronotus echinatus (Dietz, 1889) Macrops (orig.) - Downie and Arnett 1996:1504-1505
          Listronotus echinodori O'Brien, 1977 - MN, OH (Downie and Arnett 1996:1503)
          Listronotus frontalis LeConte, 1876 - IN, IL, MN, OH, ON, WI (Downie and Arnett 1996:1503)
          Listronotus grypidiodes (Dietz, 1889) Macrops (orig.) - IN, MN (Downie and Arnett 1996:1504)
          Listronotus humilus (Gyllenhal, 1834) Listroderes (orig.) - Downie and Arnett 1996:1506
          Listronotus laramiensis (Angell, 1893) Macrops (orig.)
          Listronotus maculicollis (Kirby, 1837) Macrops (orig.) - Downie and Arnett 1996:1505
          Listronotus montanus (Dietz, 1889) Macrops (orig.) - IN, IL, MN
          Listronotus sparsus (Say, 1831) Listroderes (orig.) - Downie and Arnett 1996:1505
          Listronotus squamiger (Say, 1831) Listroderes (orig.)- Downie and Arnett 1996:1502
          Listronotus tuberosus LeConte, 1876 - Downie and Arnett 1996:1502
          Listronotus vitticollis (Kirby, 1837) Macrops (orig.) - ON, south (Downie and Arnett 1996:1504)

        Lixellus LeConte, 1876 (Subfamily Cylydrorhininae)
          Lixellus filiformis LeConte, 1876 - IN, MN (Downie and Arnett 1996:1506)
          Lixellus hubbardi (LeConte, 1876) Anchodemus (orig.) - Downie and Arnett 1996:1506
          Lixellus lutulentus (Boheman, 1843) Erirhinus (orig.) - Downie and Arnett 1996:1506

        Lixus Fabricius, 1801 (Subfamily Cleoninae)
          Lixus caudifer LeConte, 1876 - IL, IN (Downie and Arnett 1996:1513)
          Lixus concavus Say, 1831 - Downie and Arnett 1996:1514
          Lixus musculus Say, 1831 - Downie and Arnett 1996:1514
          Lixus parcus LeConte, 1876 - IL, IN, ON (Downie and Arnett 1996:1514)
          Lixus punctinasus LeConte, 1876 - IL, OH, WI (Downie and Arnett 1996:1514)
          Lixus rubellus Randall, 1838 - Downie and Arnett 1996:1513
          Lixus sobrinus Casey - IL, IN, WI (Downie and Arnett 1996:1515)

        Notiodes Schönherr, 1838 (Subfamily Erirhininae)
          Notiodes ovalis (LeConte, 1876) Endalus (orig.) - IN, OH, ON, MN (Downie and Arnett 1996:1538)
          Notiodes cribricollis (LeConte, 1876) Endalus (orig.) - widespread (Downie and Arnett 1996:1538)
          Notiodes limatulus (Gyllenhal, 1836) Notiophilus (orig.) - Downie and Arnett 1996:1538

        Onychylis LeConte, 1876 (Subfamily Erirhininae)
          Onychylis longulus LeConte, 1876 - Downie and Arnett 1996:1539
          Onychylis nigrirostris (Boheman, 1843) Notiodes (orig.) - Downie and Arnett 1996; 1539

        Phytobius Schönherr, 1833 (Subfamily Ceutorhynchinae)
          Phytobius leucogaster (Marsham, 1802) Curculio (orig.) - Downie and Arnett 1996 (introduced)

        Rhinoncus Schönherr, 1825 (Subfamily Ceutorhynchinae)
          Rhinoncus asperulus (Dietz, 1896) Pelenomus (orig.) - Downie and Arnett 1996
          Rhinoncus fuliginosus (Dietz, 1896) Mecopeltus (orig.) - IN (widespread)
          Rhinoncus gracilipes (Dietz, 1896) Pelenomus (orig.) - widespread
          Rhinoncus squamosus (LeConte, 1876) Pelenomus (orig.) - Downie and Arnett 1996
          Rhinoncus sulcicollis (Fĺhraeus, 1843) Phytobius (orig.) - Downie and Arnett 1996

        Stenopelmus Schönherr, 1836 (Subfamily Erirhininae)
          Stenopelmus rufinasus Gyllenhal, 1836 - IN (widespread; Downie and Arnett 1996:1539)

        Tanysphyrus Germar, 1817 (Subfamily Erirhininae) - on Lemna sp. (duckweed)
          Tanysphyrus lemnae (Fabricius) - Downie and Arnett 1996
          Tanysphyrus ater Blatchley - Downie and Arnett 1996

    DRYOPIDAE Erichson, 1857 (Dryopid Beetles)

      Helichus Erichson, 1847
        Helichus basalis LeConte, 1852 - NMNH (web page database); see also Musgrave 1935:140-141
        Helichus fastigiatus (Say, 1824) - UMMZ (Washtenaw Co., Ann Arbor, 20-July-1901, Mus. Cat. 1634, det./ver. FNYoung)
        Helichus lithophilus (Germar, 1824) - UMMZ (Washtenaw Co., Ann Arbor, 04-Aug-1916, det. H. Nelson 1973; Oakland Co., 04-July-1909, det. H. Nelson), NMNH (web page database); see also Musgrave 1935:140
        Helichus striatus LeConte, 1852 - UMMZ (Cheboygan Co., Cheboygan, 08-Jun-1927, Sherman Moore 1029, det. FNYoung; Livingston Co., ES George Reserve, 855b & 833a coll. IJCantrall & FNYoung), NMNH (web page database); see also Musgrave 1935:142-143

      Pelonomus Erichson, 1847
        Pelonomus obscurus LeConte - IL, IN, ON (widespread central/east)

    ELMIDAE Westwood, 1838 (Riffle Beetles)

    Adult Macronychus glabratus (Copyright © 2005 Ethan Bright)
    Larva of Dubiraphia sp. (Copyright © 2005 Ethan Bright)
      Ancyronyx Erichson 1847 (Subfamily Elminae, Tribe Ancyronychini)
        Ancyronyx variegatus (Germar, 1824) Macronychus (orig.) - Brown 1983:3; Bright, pers. obs., Rouge River, Wayne Co.
          (syn.) Elmis cincta Say, 1825
          (syn.) Hydrochus parumoculatus Hardy, 1852

      Dubiraphia Sanderson 1954 (Subfamily Elminae, Tribe Elmini)
        Dubiraphia bivittata (LeConte, 1852) Elmis (orig.) - see also Brown 1983:4
        Dubiraphia minima Hilsenhoff, 1973 - Yanoviak and McCafferty 1996:205; also IN, OH, WI (Brown 1983:4)
        Dubiraphia quadrinotata (Say, 1825) Elmis (orig.) - UMMZ (Rice Creek, Calhoun Co., 01-Aug-1949, leg. V.S. Pratt, det. Sanderson, MW 1950), NMNH (web page database); see also Brown 1983:4
        Dubiraphia robusta Hilsenhoff, 1973 - WI, expect in w. UP (Hilsenhoff 1973:59; Brown 1983:4)
        Dubiraphia vittata (Melsheimer, 1844) Elmis (orig.) - Brown 1983:5; NMNH (web page database)

      Macronychus Müller 1806 (Subfamily Elminae, Tribe Macronychini)
        Macronychus glabratus Say, 1825 - UMMZ; Bright coll.; Brown 1983:12; Yanoviak and McCafferty 1996:205
          (syn.) Macronychus lateralis Melsheimer, 1844

      Microcylloepus Hinton 1935 (Subfamily Elminae, Tribe Elmini)
        Microcylloepus pusillus (LeConte, 1852) Stenelmis (orig.) - ON, WI, expect in w. UP (Brown 1983:6)

      Optioservus Sanderson 1954 (Subfamily Elminae, Tribe Elmini)
        Optioservus fastiditus (LeConte, 1850) Limnius (orig.) - Brown 1983:8; Yanoviak and McCafferty 1996:205; UMMZ (Rice Creek, Calhoun Co., 01-Aug-1949, leg. V.S. Pratt, det. Sanderson, MW 1950; Spring Brook, Kalamazoo Co., det. Sanderson, MW 1950)
        Optioservus trivittatus (Brown, 1930) Limnius (orig.) - Brown 1983:9; Yanoviak and McCafferty 1996:205; UMMZ (Rice Creek, Calhoun Co., 01-Aug-1949, leg. V.S. Pratt, det. Sanderson, MW 1950)

      Stenelmis Dufour 1835 (Subfamily Elminae, Tribe Elmini)
        Stenelmis antennalis Sanderson, 1938 - MN, WI
        Stenelmis bicarinata LeConte, 1852
          (syn.) Stenelmis maerkelii Motschulsky, 1854
          (syn.) Stenelmis vittipennis Zimmerman, 1869
        Stenelmis cheryl Brown, 1987
        Stenelmis crenata (Say, 1824) Elmis (orig.) - Brown 1983:10; Yanoviak and McCafferty 1996:205
          (syn.) Stenelmis sordida Motschulsky, 1859
        Stenelmis decorata Sanderson, 1938 - Schmude 1992
        Stenelmis douglasensis Sanderson, 1938 - Brown 1983:10 (holotype from Douglas Lake, Cheboygan Co.)
        Stenelmis fuscata Blatchley, 1925 - WI
        Stenelmis grossa Sanderson, 1938 - Schmude 1992
        Stenelmis lignicola Schmude and Brown, 1992 - Kalamazoo Co., Gull Lake Biological Station, specimen at Cook Collection - Schumde and Brown 1992
          (syn.) Stenelmis convexula Brown, 1956
          (syn.) Stenelmis markelii [sic] Finni and Skinner, 1975
          (syn.) Stenelmis crenata, S. sulcata Patrick, 1961
          (syn.) Stenelmis bicarinata, S. convexula, S. crenata, S. decorata (in part), S. hungerfordi (in part), S. sulcata Patrick et al., 1967
          (syn.) Stenelmis spp. Benke, 1984
        Stenelmis mera Sanderson, 1938 - Schmude 1992
        Stenelmis musgravei Sanderson, 1938 - see also Brown 1983:11
        Stenelmis quadrimaculata Horn, 1870 - Brown 1983:11
          (syn.) Stenelmis sulcata Blatchley, 1910
          (syn.) Stenelmis blatchleyi Musgrave, 1933
        Stenelmis sandersoni Musgrave, 1940
          (syn.) Stenelmis tarsalis Sanderson, 1938
        Stenelmis sexlineata Sanderson, 1938 - Schmude 1992

    HYDRAENIDAE Mulsant, 1844 (Minute Moss Beetles)
    The information contained on this list is tentative, based on few observations of museum specimens (currently being assessed) and literature citations (Perkins 1980).

      Gymnochthebius d'Orchymont, 1943
        Gymnochthebius nitidus (LeConte, 1850) - Perkins 1980:274-277; NMNH (web page database); holotype specimen from Eagle Harbor, Keweenaw County
          (orig.) Ochtebius nitidus LeConte, 1850:217

      Hydraena Kugelann, 1794
        Hydraena angulicollis Notman, 1921 - Perkins 1980:81-83, 443
        Hydraena pennsylvanica Kiessenwetter, 1849 - Perkins 1980:81, 96-98, 443; neootype from Bryant Road, Emmet County; record from Grand Ledge, Eaton Co. as H. atlantica (Perkins 1980)
          (syn.) Hydraena atlantica Perkins, 1980
      Ochthebius Leach, 1815
        Ochthebius (Asiobates) cribricollis LeConte, 1850 - Perkins 1980:386, 388-389, lectotype from Eagle Harbor, Keweenaw County
        Ochthebius (Ochtebius) kaszabi Janssens, 1967 - MN, ON (may be found in Isle Royale)(Perkins 1980:282, 325-328)
        Ochthebius (Ochtebius) lineatus LeConte, 1852 - WI, western, but one record from northern WI, expect in w UP (Perkins 1980:314-318, 323)
          (syn.) Ochthebius milleri Hatch, 1965

    HYDROPHILIDAE Latreille, 1802 (Water Scavenger Beetles)
    The information contained on this list is tentative, based on few observations of museum specimens (currently being assessed) and a limited literature search (Downie and Arnett 1996, Willson 1970). Terrestrial species of Sphaeridiinae, including those in the genera Cercyon, Cryptopleurum, Paraceryon, Pemelus, Phaenonotum, Sphaeridium, and Tectosternum are not included, as they are associated with decaying plant or fungal matter, and animal dung, and are only rarely (or incidentally) collected in aquatic systems. A number of species of Cercyon and Phaenonotum are eith semi- or peri-aquatic species, and associated with aquatic habitats, and these are included. Consult Smetana (1988) or Downie and Arnett (1996) for more information.

      Anacaena Thomson, 1859 (Subfamily Hydrobiinae, Tribe Hydrobiini)
        Anacaena lutescens Stephens, 1829 - (as A. limbata (Fabricius, 1792), see Hilsenhoff 1995c), UMMZ; see also Willson 1970:80-81; Smetana 1988:87-88

      Berosus Leach, 1817 (Subfamily Berosinae)
        Berosus aculeatus LeConte, 1855 - Smetana 1988:51; Downie and Arnett 1996
        Berosus fraternus LeConte, 1855 - Downie and Arnett 1996, NMNH (web page database); see also Smetana 1988:54-55
          (syn.) Berosus californicus Motschulsky, 1859
        Berosus hatchi Miller, 1965 - ON, WI (Smetana 1988:53-54)
        Berosus infuscatus LeConte, 1855 - Charity Island (44.030°N, 83.437°W), specimens in UMMZ, NMNH (web page database)
        Berosus ordinatus LeConte - IN, MN (and eastward)
        Berosus pantherinus LeConte, 1855 - IL, IN, OH, ON, WI
        Berosus peregrinus (Herbst, 1797) Hydrophilus (orig.) - UMMZ; see also Smetana 1988:50
          (syn.) Berosus auritus Melsheimer, 1846
          (syn.) Berosus peregrinus, LeConte 1855, Matta, 1974
          (syn.) Berosus subsignatus LeConte, 1855
          (syn.) Berosus immaculatus Zimmermann, 1869
        Berosus striatus (Say, 1825) Hydrophilus (orig.) - UMMZ, NMNH (web page database); see also Smetana 1988:51-52
        Berosus stylifer Horn, 1873 - Smetana 1988:55-56
          (syn.) Berosus falcatus Sharp, 1882

      Cercyon Leach, 1817 (Subfamily Spaheridiinae, Tribe Cercyonini)
        Cercyon (Prostercyon) roseni Knisch, 1922 - Smetana 1988:136-137 (peri- to semi-aquatic, along wetlands, ponds, bogs and lakes)
        Cercyon (Cercyon) connivens Fall, 1924 - Smetana 1988: 141 (peri-aquatic)
        Cercyon (Cercyon) herceus Smetana, 1978 - Smetana 1988: 142-143 (peri-aquatic)
          (syn.) Cercyon tristis, Horn 1890 (nec Illiger, 1801)
          (syn.) Cercyon herceus Smetana, 1978; plus spp. herceus and frigidus
        Cercyon (Cercyon) erraticus Smetana, 1978 - Smetana 1988: 144 (peri-aquatic)
        Cercyon (Cercyon) mendax Smetana, 1978 - Smetana 1988: 145 (peri-aquatic)
        Cercyon (Cercyon) praetextatus (Say, 1825) Sphaeridium (orig.) - Smetana 1988: 145-146
          (syn.) Cercyon testaceus Blatchley, 1923
        Cercyon (Cercyon) marinus Thomson, 1853 - Smetana 1988: 147-148 (semi-aquatic)
        Cercyon (Cercyon) cinctus Smetana, 1978 - Smetana 1988: 148 (peri-aquatic, similar habitat preferences with marinus)

      Chaetarthria Stephens, 1833 (Subfamily Chaetarthriinae)
        Chaetarthria atra (LeConte, 1863) Cyllidium (orig.) - IL, IN (Smetana 1988: 57-58)
          (syn.) Chaetarthria nigrella, Wickham 1895 (nec LeConte, 1861)
        Chaetarthria pallida (LeConte, 1861) Cyllidium (orig.) - Cheboygan Co., UMMZ, NMNH (web page database); see also Smetana 1988: 58
          (syn.) Cyllidum nigriceps LeConte, 1861

      Crenitis Bedel, 1881 (Subfamily Hydrobiinae)
        Crenitis digesta (LeConte, 1855) Hydrobius (orig.) - Willson 1970: 78-79; see also Smetana 1988: 81-82)
          (syn.) Paracymus digestus Winters, 1926
          (syn.) Crenophilus digestus d'Orchymont, 1942

      Cymbiodyta Bedel, 1881 (Subfamily Hydrobiinae, Tribe Helocharini)
        Cymbiodyta acuminata Fall, 1924 - UMMZ; Willson 1970: 52-53; Smetana 1974: 68-70; see also Smetana 1988: 110-111
        Cymbiodyta blanchardi Horn, 1890 - UMMZ; Willson 1970: 53; see also Smetana 1974: 61-64; Smetana 1988: 107-108
        Cymbiodyta chamberlaini Smetana, 1974 - UMMZ; Smetana 1974: 57-61; Smetana 1988: 106-107
        Cymbiodyta minima Notman, 1919 - UMMZ, NMNH (web page database); Willson 1970: 54; Smetana 1974: 73-76; see also Smetana 1988: 111-112
          (syn.) ?Hydrobius marginellus Kirby, 1837, in Richardson
          (syn.) Philidrus lacustris Horn, 1873
          (syn.) Cymbiodyta lacustris Wickham, 1895
          (syn.) Cymbiodyta acuminata Hatch, 1933 (misid).
        Cymbiodyta semistriata (Zimmermann, 1869) Philhydrus (orig.) - UMMZ, NMNH (web page database); Smetana 1974: 48-51; see also Smetana 1988: 103, 105
          (syn.) Philhydrus fimbriatus LeConte, 1855
          (syn.) Philidrus fimbriatus Provancher, 1877
          (syn.) Cymbiodyta fimbriata, Horn 1890 and authors Cymbiodyta toddi Spangler, 1966 - Willson 1970: 56; Smetana 1974: 51-54; see also Smetana 1988: 104, 106
          Cymbiodyta vindicata Fall, 1924 - UMMZ, NMNH (web page database); Willson 1970: 55-56; Smetana 1974: 64-67; see also Smetana 1988: 108-109

        Dibolocelus Bedel, 1892 (Subfamily Hydrophilinae)
          Dibolocelus ovatus (Gemminger & Harold, 1868) Hydrophilus (orig.) - Willson 1970; see also Smetana 1988: 124
            (syn.) Hydrophilus ovilis Ziegler, 1844 (nec Laport, 1840)

        Enochrus Thomson, 1859 (Subfamily Hydrobiinae, Tribe Helocharini)
          Enochrus (Methydrus) blatchleyi (Fall, 1924) Philhydrus (orig.) - Willson 1970: 62; see also Smetana 1988: 92
          Enochrus (Methydrus) cinctus (Say, 1824) Hydrophilus - Willson 1970: 63-64; record at Albertson College of Idaho Collection; see also Smetana 1988: 92-93
            (syn.) Philhydrus limbalis Melsheimer, 1846
          Enochrus collinus (Fall, 1924) Philhydrus (orig.) - WI
          Enochrus (Methydrus) consors LeConte, 1863 Philhydrus (orig.) - Willson 1970: 65; see also Smetana 1988: 93)
          Enochrus (Methydrus) consortus Green, 1946 - Willson 1970: 64-65; see also Smetana 1988: 93-94
          Enochrus (Lumetus) diffusus (LeConte, 1855) Philhydrus (orig.) - record at Albertson College of Idaho Collection; western species with isolated eastern populations (Smetana 1988: 99); see also Willson 1970: 66-67
          Enochrus (Lumetus) hamiltoni (Horn, 1890) Philhydrus (orig.) - Downie and Arnett 1996; see also Smetana 1988: 99-100
            (syn.) Philhydrus lividus Walker, 1866
            (syn.) Philhydrus conjunctus Fall, 1901
            (syn.) Enochrus collinus Brown, 1931
            (syn.) Enochrus horni Leech, 1950
            (syn.) Enochrus pacificus Leech, 1950
            (syn.) Enochrus pyretus Leech, 1950
          Enochrus (Methydrus) ochraceus (Melsheimer, 1846) Philhydrus (orig.) - Willson 1970: 62-63; record at Albertson College of Idaho Collection; see also Smetana 1988: 95
            (syn.) Philhydrus lacustris LeConte, 1855
            (syn.) Philhydrus simplex LeConte, 1863
          Enochrus (Methydrus) perplexus (LeConte, 1855) Philhydrus (orig.) - Willson 1970: 54-55 (as C. fimbriata), 65-66; see also Smetana 1988: 94
            (syn.) Cymbiodyta fimbriatua (Melsheimer, 1846)
          Enochrus (Methydrus) pygmaeus nebulosus (Say, 1824) Hydrophilus - Willson 1970: 60-61; see also Smetana 1988: 96
            (syn.) Hydrophilus pygmaeus Fabricius, 1792
          Enochrus (Methydrus) sayi Gunderson, 1977 - Downie and Arnett 1996; Smetana 1988: 97

        Helocombus Horn, 1890 (Subfamily Hydrobiinae, Tribe Helocharini)
          Helocombus bifidus (LeConte, 1855) Philhydrus (orig.) - Willson 1970: 50-51; see also Smetana 1988: 113

        Helophorus Fabricius, 1775 (Subfamily Helophorinae)
          Helophorus (Rhopalelophorus) angusticollis d'Orchymont - Downie and Arnett 1996; see also Smetana 1988: 31, 43-44
          Helophorus (Rhopalelophorus) artus Smetana - IL
          Helophorus (Rhopalelophorus) furius Smetana, 1987 - ON, WI (northern: Smetana 1988: 27-28)
            (syn.) Helophorus frater Smetana, 1985 (nec d'Orchymont, 1926)
          Helophorus (Helophorus) grandis Illiger, 1798 Elophorus (orig.) - record from Huron Co.; see Smetana 1988: 20-21 (introduced species from Europe)
            (syn.) Helophorus aquaticus, Darlington 1927, Brown 1940 (nec Linné 1758)
          Helophorus (Rhopalelophorus) inflectus McCorkle, 1965 Elophorus (orig.) - Smetana 1988: 25-26; Downie and Arnett 1996
          Helophorus (Rhopalelophorus) lacustris LeConte, 1850 - Downie and Arnett 1996; see also Smetana 1988: 39-40
          Helophorus (Rhopalelophorus) latipenis - WI (syn. or ssp.?)
          Helophorus (Rhopalelophorus) linearis LeConte, 1850 - IL, IN (Waltz 2002: 178), OH, ON, WI (Smetana 1988: 36-37)
            (syn.) Helophorus pallens Casey, 1884
            (syn.) Helophorus oblitus Sharp, 1915
          Helophorus (Rhopalelophorus) lineatus Say, 1823 - Downie and Arnett 1996; see also Smetana 1988: 29-30
            (syn.) Helophorus ventralis Motschulsky, 1860
            (syn.) Helophorus obsoletesulcatus Motschulsky, 1860
          Helophorus (Rhopalelophorus) marginicollis Smetana, 1985 - Smetana 1988: 44; Downie and Arnett 1996
          Helophorus (Rhopalelophorus) nitiduloides d'Orchymont, 1945 - Downie and Arnett 1996; see also Smetana 1988: 37-38
          Helophorus (Rhopalelophorus) nitidulus LeConte, 1855 - Smetana 1988: 28-29; Downie and Arnett 1996
            (syn.) Helophorus gregalis d'Orchymont, 1945
          Helophorus (Rhopalelophorus) oblongus LeConte, 1850 - Downie and Arnett 1996; see also Smetana 1988: 24-25
          Helophorus (Rhopalelophorus) orchymonti Smetana, 1985 - ON, WI (Smetana 1988: 43)
          Helophorus (Rhopalelophorus) orientalis Motschulsky, 1860 - Downie and Arnett 1996; see also Smetana 1988: 34-35
            (syn.) Helophorus brevipalpis, McCorkle, 1965 (nec Bedel, 1881)
          Helophorus (Rhopalelophorus) sempervarians Angus, 1970 - Downie and Arnett 1996; see also Smetana 1988: 32-33
            (syn.) Elophorus jacutus, McCorkle, 1965 (nec Poppius, 1907)
            (syn.) Helophorus jacutus, d'Orchymont, 1945 (nec Poppius, 1907)
          Helophorus (Cyphelophorus) tuberculatus Gyllenhal - Downie and Arnett 1996; see also Smetana 1988: 45-46

        Hydrobius Leach, 1815 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Hydrobius fuscipes (Linnaeus, 1758) Dytiscus (orig.) - Willson 1970: 69, NMNH (web page database), record at Albertson College of Idaho Collection; see also Smetana 1988: 69-70
          Hydrobius melaenus (Germar, 1824) Sphaeridium (orig.) - Willson 1970: 69-71 (as H. globosus); see also Smetana 1988: 70-71
            (syn.) Hydrophilus globosus Say, 1824
            (syn.) Hydrobius globosus LeConte, 1855
            (syn.) Hydrobius globulosus Provancher, 1877
            (syn.) Hydrobius melaenus Winters, 1926
          Hydrobius tumidus LeConte, 1855 - IN

        Hydrochara Berthold, 1827 (Subfamily Hydrophilinae)
          Hydrochara leechi Smetana, 1980 - IL, OH, WI, may occur in s LP (Smetana 1980: 38-41)
          Hydrochara obtusata (Say, 1823) Hydrophilus - Willson 1970: 22-23; NMNH (web page database); Smetana 1980: 24-30; see also Smetana 1988: 127
          Hydrochara simula Hilsenhoff & Tracy - Hilsenhoff and Tracy 1982: 101; see also Smetana 1988: 128
          Hydrochara soror Smetana, 1980 - Smetana 1980: 30-34; see also Smetana 1988: 128
          Hydrochara spangleri Smetana, 1980 - IL, IN, OH, WI, may occur in sw LP (Smetana 1980: 34-36)

        Hydrochus Leach, 1817 (Subfamily Hydrochinae)
          Hydrochus brevitarsis Knisch, 1922 - Smetana 1988: 12; Hilsenhoff 1995: 42
          Hydrochus currani Brown, 1929 - ON, WI (Smetana 1988: 11-12)
          Hydrochus granulatus Blatchley - IN, ON, WI
          Hydrochus excavatum LeConte - IN
          Hydrochus granulatus Blatchly, 1910 - QB-BC, NY-MN (Smetana 1988: 17-18)
            (syn.) Hydrochus ouelleti Leech, 1848
          Hydrochus inaequalis LeConte - IN
          Hydrochus neosquamifer Smetana, 1988 - ON, WI (Smetana 1988: 14-15)
          Hydrochus pseudosquamifer Miller, 1965 - IN, ON, WI (Smetana 1988: 15)
          Hydrochus rufipes Melsheimer - IL, IN, OH, WI (Smetana 1988: 18)
            (syn.) Hydrochus impressus Zimmermann, 1868
          Hydrochus rugosus Mulsant - IN
          Hydrochus scabratus Mulsant, 1844 - Downie and Arnett 1996; see also Smetana 1988: 11
            (syn.) Hydrochus gibbosus Melsheimer, 1846
          Hydrochus squamifer LeConte, 1855 - IN, ON, WI (Smetana 1988: 13-14)
            (syn.) Hydrochus laticollis Notman, 1919
          Hydrochus subcupreous Randall, 1838 - IN, WI (Smetana 1988: 15-17)

        Hydrophilus Geoffroy, 1762 (Subfamily Hydrophilinae)
          Hydrophilus triangularis Say, 1823 - Willson 1970: 24-25; see also Smetana 1988: 123-124

        Laccobius Erichson, 1837 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Laccobius agilis Randall, 1838 Hydrophilus (orig.) - Willson 1970: 38-39; see also Smetana 1988: 61-62
            (syn.) Laccobius ellipticus, Stace Smith 1930 (nec LeConte, 1855)
            (syn.) Laccobius agilis Gentili, 1985
          Laccobius carri d'Orchymont, 1942 - ON, may occur in UP (Smetana 1988: 67-68)
          Laccobius cinereus columbianus Miller, 1965 - Downie and Arnett 1996; see also Smetana 1988: 64-65
          Laccobius fuscipunctatus Hilsenhoff, 1995 - WI (Hilsenhoff 1995c: 112-114)
          Laccobius minutoides d'Orchymont, 1942 - Willson 1970: 39-40; see also Smetana 1988: 62-63
          Laccobius reflexipennis Cheary, 1971 - ON, WI (Smetana 1988: 66-67)
            (syn.) Laccobius reflexipennis Malcolm, 1979
          Laccobius spangleri Cheary, 1971 - Willson 1970: 40-44; Cheary 1971; NMNH (web page database); see also Smetana 1988: 67
            (inv.) Laccobius spangleri Willson, 1970, manuscript, never published
            (syn.) Laccobius spangleri Malcolm, 1979
          Laccobius teneralis Cheary, 1971 - ON, may occur in s. LP (Smetana 1988: 65-66)
          Laccobius truncatipenis Miller, 1965 - Willson 1970: 44-48 (as L. arenarius); Cheary 1971 (as L. arenarius); NMNH (web page database); see also Smetana 1988: 68-69
            (syn.) Laccobius arenarius Willson 1970, unpublished MS thesis
            (syn.) Laccobius arenarius Cheary, 1971

        Paracymus Thomson, 1867 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Paracymus confluens Wooldridge, 1966 - Wooldridge 1966; Willson 1970: 74-75; Smetana 1988: 75-76
          Paracymus confusus Wooldridge, 1966 - Downie and Arnett 1996; see also Willson 1970: 75; Smetana 1988: 77
          Paracymus despectus (LeConte, 1863) Hydrobius - Willson 1970: 73-74; NMNH (web page database); see also Smetana 1988: 78
          Paracymus subcupreus (Say, 1825) Hydrobius - Willson 1970: 75-76, NMNH (web page database); see also Smetana 1988: 76-77

        Phaenonotum Sharp, 1882 (Subfamily Spaheridiinae, Tribe Sphaeridiini)
          Phaenonotum exstriatum (Say, 1835) Hydrophilus (orig.) - Willson 1970: 13-14; Smetana 1988: 130 (often semi-aquatic, at edges of wet habitats and water, wet moss and Sphagnum)
            (comb.)Cyclonotum estriatum LeConte, 1855 (unjustified emendation)
            (syn.) Cyclonotum semiglobosum Zimmermann, 1869
            (syn.) Phaenonotum dubium Sharp, 1882
            (spell.) Phaenonotum estriatum (Say, 1835), Willson 1970: 13

        Sperchopsis LeConte, 1862 (Subfamily Hydrobiinae, Tribe Hydrobiini)
          Sperchopsis tessellatus (Ziegler, 1844) Spercheus (orig.) - Willson 1970: 71-72; record at Albertson College of Idaho Collection; Bright coll.; see also Smetana 1988: 72
            (syn.) Sperchopsis tessellatus LeConte, 1861
            (syn.) Hydrobius tessellatus Horn, 1873
            (syn.) Hydrocyclus tessellatus Winters, 1926
            (syn.) Sperchopsis tessellatus Spangler, 1961

        Tropisternus Solier, 1834 (Subfamily Hydrophilinae)
          Tropisternus blatchleyi modestus d'Orchymont, 1938 - Willson 1970: 32 (as T. modestus); see also Smetana 1988: 119-120
            (syn.) Tropisternus modestus d'Orchymont, 1938
          Tropisternus columbianus Brown, 1931 - Willson 1970: 29-30; see also Smetana 1988: 122
          Tropisternus ellipticus LeConte, 1855 Hydrophilus - IL, WI (Smetana 1988: 118-119; see also Willson 1970: 29)
            (syn.) Tropisternus affinis Motschulsky, 1859
            (syn.) Tropisternus concolor Sharp, 1882
          Tropisternus glaber (Herbst, 1797) Hydrophilus (orig.) - Willson 1970: 33-34; Ryker 1972; see also Smetana 1988: 121
          Tropisternus lateralis nimbatus (Say, 1823) Hydrophilus (orig.) - Willson 1970: 28; Ryker 1972 (as T. nimbatus); see also Smetana 1988: 118
            (syn.) Tropisternus nimbatus Summers, 1874
            (syn.) Tropisternus lateralis Horn, 1876; LaRivers 1949
            (syn.) Tropisternus lateralis nimbatus Young, 1954
          Tropisternus mixtus (LeConte, 1855) Hydrophilus (orig.) - Willson 1970: 30-31; Ryker 1972; see also Smetana 1988: 119
          Tropisternus natator d'Orchymont, 1938 - Willson 1970: 32-33; Ryker 1972; see also Smetana 1988: 120-121
            (syn.) Tropisternus glaber, LeConte, 1859; Wooldridge, 1967
            (syn.) Hydrophilus glaber, LeConte, 1855 (nec Herbst, 1797)

      LAMPYRIDAE Latreille, 1817 (Firefly Beetles)

      Several taxa of these fascinating beetles have larvae known to live in the air spaces of aquatic plants, or are associated with aquatic or semiaquatic plants in aquatic habitats. A number of species have larvae that are adapted to feed on aquatic snails, and perhaps annelids; some larvae of Asian species have tracheal gills that permit a truly aquatic existence. There are 6 species in 2 genera that have been recorded in Michigan, with another 3 species in 2 genera likely to be found based on their regional distribution. This list is based on information kindly provided by Dr. Jim Lloyd (pers.comm. 2003), and records listed in Downie and Arnett (1996).

        Photinus LaPorte, 1833

          Photinus ardens LeConte, 1851 - Downie and Arnett 1996; Lloyd pers.comm. (Marquette, Delta, Chippewa, Charlevoix, Cheboygan, Roscommon, Huron, St. Clair, Washtenaw)
          Photinus consimilis Green (complex), 1956 - Downie and Arnett 1996; Lloyd pers.comm., as consimilis-complex (Schoolcraft, Chippewa, Oceana, Livingston, Oakland, St. Clair)
          Photinus obscurellus LeConte, 1851 - Downie and Arnett 1996; Lloyd pers.comm (Houghton, Iron, Dickinson, Marquette, Mackinaw, Chippewa, Berrien, St. Joseph, Washtenaw, Wayne, St. Clair)

        Pyractomena LeConte, 1850

          Pyractomena angulata (Say, 1825) Lampyris (orig.) - e. USA, Canada (Lloyd, pers.comm.; Luk et al. 2011: 68, 72)
          Pyractomena borealis (Randall, 18258 Lampyris (orig.) - OH, e. USA, Canada (Archangelsky and Branham 1998; Lloyd, pers.comm.; Luk et al. 2011: 69, 73)
          Pyractomena dispersa Green, 1958 - Downie and Arnett 1996; Lloyd pers. comm. (Kent, Washtenaw, Wayne)
          Pyractomena linearis LeConte, 1851 - Downie and Arnett 1996; Lloyd pers.comm. (Marquette, Dickinson, Mackinac, Livingston, Washtenaw)
          Pyractomena lucifera Melsheimer, 1846 - Downie and Arnett 1996; Lloyd pers.comm. (Midland, Clinton, Livingston, Washtenaw, Oakland)
          Pyractomena sinuata Green, 1958 - IL, MN, WI (Lloyd pers.comm.)

        Pyropyga Motschulsky, 1852

          Pyropyga decipiens (Harris, 1836) Lampyris (orig.) - Lloyd pers.comm. (Keweenaw, widespread in LP)
          Pyropyga nigricans (Say, 1823) Lampyris (orig.) - Lloyd 1999; Lloyd pers.comm. (widespread in state)

      LUTROCHIDAE Kasap & Crowson, 1975 (Long-toed or Travertine Beetles) (Polyphaga)
      Lutrochus luteus adult - from Brown 1972

      These beetles share the same habitats as that of dryopid and elmid riffle beetles (Dryopidae, Elmidae) and water-pennies (Psephenidae), although there is a preference for calcarious deposits from mineral springs (Brown 1972) (or possibly areas with high rates of instream photosynthetic activity, when calcium carbonate precipitates). There is one species that reaches our area. In Wisconsin, larvae were found in rocks, and adults were found in various lotic habitats of mixed substrate, with ovipositing adults occuring on wood and the travertine of rocks in shallow riffles (Hilsenhoff and Schmude 1992).

        Lutrochus Erichson, 1847
          Lutrochus laticeps Casey, 1893 - Brown 1972

      PSEPHENIDAE Lacordaire, 1854 (Water Pennies)

      Larva of Psephenus herricki (Copyright © 2006 Ethan Bright)
      Psephenus herricki - larva - copyright 2006 Ethan Bright

        Ectopria LeConte, 1853 (Subfamily Eubriinae)
          Ectopria leechi Brigham, 1981
          Ectopria nervosa (Melsheimer, 1845) Eubria (orig.) - Yankoviak and McCafferty 1996: 205
            (syn.) Eubria thoracica Ziegler, 1845
            (syn.) Ectopria tarsalis LeConte, 1853
            (syn.) Ectopria tibialis LeConte, 1853

        Psephenus Haldeman 1853 (Subfamily Psepheninae)
          Psephenus herricki (De Kay, 1844) Fluvicola (orig., described as an isopod) - Brown and Murvosh 1974: 315-322; widespread in state
            (nomen nudum) Eurypalpus lecontei De Jean, 1837
            (syn.) Psephenus lecontei Haldeman, 1853

      PTILODACTYLIDAE Laporte, 1836 (Toad-Winged Beetles)

      Larvae are known to live in semiaquatic or aquatic habitats, such as decaying damp logs and organic muck. Recently the larva of Paralichas (=Odontyonyx) trivattis has been described, reared from specimens sampled from spring seeps in eastern Pennsylvania (Funk and Fenstermacher 2002). Downie and Arnett (1996) list a questionable record from Indiana, and since the biology and distribution of this species are not well known, I have included this taxa as it conceivably may occur in our area, especially in morainal/outwash areas of southern Michigan where groundwater seeps occur.

        Odontonyx Guérin-Méneville, 1843

          Odontonyx trivittis (Germar, 1824) Dasytes - IN?, PA

      SCIRTIDAE Fleming, 1821 (=Helodidae) (Marsh Beetles)

        Cyphon Paykull, 1799
          Cyphon collaris (Guérin-Meneville, 1843) Elodes (orig.) - IN, ON
          Cyphon confusus Brown, 1930 - Young and Stribling 1990: 201-202
            (syn.) Cyphon horioni Klausnitzer, 1976
          Cyphon drymophilous Young & Stribling, 1990 - OH, WI (Young and Stribling 1990: 202-203)
          Cyphon obscurus (Guérin-Meneville, 1843) Elodes (orig.) - IN, OH, ON
          Cyphon padi (Linnaeus, 1758) Chrysomela (orig.) - IN, ON
          Cyphon ruficollis (Say, 1825) Lampyris (orig.) - IN (widespread east and central)
          Cyphon variabilis (Thunberg, 1785) Cantharis - Downie and Arnett 1996

        Elodes Latreille, 1796

          Elodes fuscipennis Guérin-Meneville 1843 - IN
          Elodes maculicollis (Horn, 1880) Helodes (orig.) - ON
          Elodes pulchella Guérin-Meneville, 1843 - IN
          Elodes thoracica Guérin-Meneville, 1843 - IN, ON

        Microcara Thomson 1859

          Microcara explanata LeConte, 1866 Helodes (orig.) - Downie and Arnett 1996, shores of Lake Michigan

        Prionocyphon Redtenbacher, 1858

          Prionocyphon discoideus (Say, 1825) Cyphon (orig.) - Downie and Arnett 1996

        Scirtes Illiger, 1807

          Scirtes orbiculatus (Fabricius, 1801) Cyphon (orig.) - IN, OH
          Scirtes tibialis (Guérin-Meneville, 1843) Scirtes (orig.) - IN, ON, WI

      STAPHYLINIDAE Latreille, 1802 (Rove Beetles)

      This large family of slender beetles are characteristic in having short, truncated elytra that leaves several abdominal segments dorsally visible. Most species are terrestrial, but a few are associated with aquatic environments, principally with shore-line environments and the surface of littoral habitats. Psephidonus is common under stones at or near the edge of running water. Carpelimus and Thinobius is generally found along littoral areas of lakes and rivers among leaf litter. Not all species of Bledius and Stenus are semiaquatic, and provisionally terrestrial taxa are included in the list based on citations in Downie and Arnett (1996); museum and other collection specimens have have only begun to be examined.

        Bledius Leach, 1819 (Subfamily Osytelinae)

          Bledius analis LeConte, 1863 - Downie and Arnett 1996
          Bledius annularis LeConte, 1863 - IN (widespread)
          Bledius armatus Say - UMMZ Insect Collection (syn. of what name?)
          Bledius assimilis Casey, 1889 - Downie and Arnett 1996
          Bledius confusus LeConte, 1877 - Downie and Arnett 1996
          Bledius emarginatus (Say, 1834) Oxytelus - Downie and Arnett 1996
          Bledius fumatus LeConte, 1863 - Downie and Arnett 1996
          Bledius gravidus Casey, 1889 - Downie and Arnett 1996
          Bledius ineptus Casey, 1889 - Downie and Arnett 1996
          Bledius nitidicollis LeConte, 1863 - IL, OH (and northwest)
          Bledius omega Herman, 1983 - Downie and Arnett 1996
          Bledius pallipennis (Say, 1823) Oxytelus - IL, OH
          Bledius rotundicollis LeConte, 1877 - IL, MN, WI
          Bledius ruficornis LeConte, 1863 - Downie and Arnett 1996
          Bledius semiferrugineus LeConte, 1863 - UMMZ Insect Collection
          Bledius sinuatus LeConte, 1877 - IL, ON
          Bledius suturalis LeConte, 1863 - ON (widespread)
          Bledius tarandus Herman, 1970 - ON, WI
            (syn.) Bledius borealis Blatchley
          Bledius tau LeConte, 1877 - Downie and Arnett 1996
          Bledius trugidus Casey, 1889 - Downie and Arnett 1996
          Bledius viriosus Herman, 1983 - ON, WI

        Carpelimus Leach, 1819 (Subfamily Osytelinae)

          Carpelimus bilineatus (Stephen), 1832 Carpalimnus [sic] - IN
          Carpelimus delicatus (Casey, 1889) Trogophloeus - Downie and Arnett 1996
          Carpelimus fallax (Casey, 1889) Trogophloeus - Downie and Arnett 1996
          Carpelimus morio (Erichson, 1840) Trogophloeus - Downie and Arnett 1996
          Carpelimus quadripunctatus (Say, 1834) Stenus - IN, OH

        Psephidonus Gistel, 1856 (=Geodromicus Redtenbacher) (Subfamily Omaliinae)

          Psephidonus brunneous (Say, 1823) Anthophagus - UMMZ Insect Division
          Psephidonus strictus (Fauvel), 1889 - Downie and Arnett 1996
          Psephidonus ovipennis (LeConte, 1878) Geodromicus - widespread, disjunct (New York, Utah)

        Stenus Latreille, 1796 (Subfamily Steninae)

          Stenus advenus (Casey, 1884) Areus - Downie and Arnett 1996
          Stenus alacer Casey, 1884 - IL, IN
          Stenus argus Gravenhorst, 1806 - IN
          Stenus angustus Casey, 1884 - Downie and Arnett 1996
            (syn.) Stenus indigens Casey, 1884
            (syn.) Stenus pettiti Casey, 1884
            (syn.) Stenus placidus Casey, 1884
            (syn.) Stenus mendax Casey, 1884
          Stenus annularis Erichson, 1840 - Downie and Arnett 1996
          Stenus brivioi Puthz, 1972 - Downie and Arnett 1996
          Stenus callosus Erichson, 1840 - ? (no locality in Downie and Arnett 1996)
          Stenus canaliculatus Gyllenhal, 1827 - Downie and Arnett 1996
          Stenus capucinus Boheman, 1858 - Downie and Arnett 1996
            (syn.) Stenus varipes Casey, 1884
          Stenus carinicollis Casey, 1884 - Downie and Arnett 1996
          Stenus caseyi Puthz, 1971 - Downie and Arnett 1996
          Stenus colonus Erichson, 1840 - Downie and Arnett 1996
            (syn.) Stenus militaris Casey, 1884
            (syn.) Stenus suspectus Blatchley
            (syn.) Stenus obstrusus Casey, 1884
            (syn.) Stenus integer Casey, 1884
          Stenus comma LeConte, 1863 - Downie and Arnett 1996
          Stenus colon Say, 1834 - Downie and Arnett 1996
          Stenus croceatus (Casey, 1884) - Downie and Arnett 1996
          Stenus curtus Casey, 1884 - Downie and Arnett 1996
          Stenus dissentiens (Casey, 1884) - Downie and Arnett 1996
            (syn.) Stenus hubbardi (Casey, 1884)
          Stenus egenulus Puthz - Downie and Arnett 1996
          Stenus egenus Erichson, 1840 - IN (widespread)
          Stenus erythropus Melsheimer, 1844 - Downie and Arnett 1996
          Stenus femoratus Say, 1834 - Downie and Arnett 1996
          Stenus flavicornis Erichson, 1840 - Downie and Arnett 1996
          Stenus fulvoguttatus Notman, 1920 - Downie and Arnett 1996
          Stenus gratiosus Casey, 1884 - Downie and Arnett 1996
            (syn.) Stenus hirsutus Casey, 1884
          Stenus immarginatus Maklin, 1853 - widespread east
            (syn.) Stenus monticola Casey, 1884
          Stenus insperatus Puthz, 1972 - Downie and Arnett 1996
          Stenus intrusus Casey, 1884 - Downie and Arnett 1996
          Stenus juno Fabricius, 1801 - Downie and Arnett 1996
          Stenus laccophilus Casey, 1884 - UMMZ insect collection
            (syn.) Stenus edax Notman
            (syn.) Stenus rugifer Casey, 1884
          Stenus mammops Casey, 1884 - Downie and Arnett 1996
          Stenus morio Gravenhorst, 1806 - Downie and Arnett 1996
            (syn.) Stenus morio Gravenhorst, 1806
          Stenus nanus Stephen, 1832 - Downie and Arnett 1996
          Stenus neglectus Casey, 1884 - Downie and Arnett 1996
          Stenus parallelopipedus Casey, 1884 - Downie and Arnett 1996
            (syn.) Stenus animatus Casey, 1884
          Stenus pauperculus Casey, 1884 - Downie and Arnett 1996
          Stenus pubescens Stephens, 1832 - Downie and Arnett 1996 (native of Palearctic?)
          Stenus pudicus Casey, 1884 - Downie and Arnett 1996
            (syn.) Stenus juvencus Casey, 1884
          Stenus pumilio Erichson, 1839 - Downie and Arnett 1996
          Stenus punctatus Erichson, 1840 - Downie and Arnett 1996
          Stenus pusillimus Puthz, 1971 - Downie and Arnett 1996
          Stenus reconditus (Casey, 1884) - Downie and Arnett 1996
            (syn.) Stenus canadensis (Casey, 1884)
          Stenus rossi Sanderson, 1957 - Downie and Arnett 1996
          Stenus scrupens Casey, 1884 - Downie and Arnett 1996
          Stenus semicolon LeConte, 1863 - Downie and Arnett 1996
          Stenus strangulatus Casey, 1884 - Downie and Arnett 1996
          Stenus scabiosus Casey, 1884 - Downie and Arnett 1996
          Stenus sibiricus Sahlberg, 1880 - Downie and Arnett 1996
            (syn.) Stenus deductor Casey, 1884
            (syn.) Stenus montanus Casey, 1884
          Stenus stygicus Say, 1834 - Downie and Arnett 1996
          Stenus tenuis Casey, 1884 - Downie and Arnett 1996
            (syn.) Stenus pauper Casey, 1884
            (syn.) Stenus subtilis Casey, 1884
            (syn.) Stenus ingratus Casey, 1884
          Stenus varipes Casey, 1884 - Downie and Arnett 1996
          Stenus vicinus Casey, 1884 - Downie and Arnett 1996
          Stenus vinnulus Casey, 1884 - Downie and Arnett 1996

        Thinobius Kiesenwetter, 1844 (Subfamily Osytelinae)

          Thinobius brachypterusLeConte, 1877 - Downie and Arnett 1996
          Thinobius fimbriatus LeConte, 1877 - Downie and Arnett 1996

References

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    Archangelsky M, Branham MA. 1998. Description of the preimaginal stages of Pyractomena borealis (Randall, 1838)(Coleoptera: Lampyridae) and notes on its biology. Proceedings of the Entomological Society of Washington 100(3):421-430).
    Askevold IS. 1991. Classification, reconstructed phylogeny, and geographic history of the new world members of Plateumaris Thomson, 1859 (Coleoptera: Chrysomelidae: Donaciinae). Memoirs of the Entomological Society of Canada 157:1-175.
    Atton FM. 1990. Gyrinus (Gyrinulus) cavatus sp. nov. from North America described and compared with Gyrinus (Gyrinulus) minutus Fabricius (Coleoptera: Gyrinidae). The Canadian Entomologist 122(4):651-657.
    Brigham WU. 1981. Ectopria leechi, a new false water penny from the United States (Coleooptera: Eubriidae). Pan-Pacific Entomologist 57:313-320.
    Brown HP. 1972. Aquatic dyropoid beetles (Coleoptera) of the United States. Biota of Freshwater Ecosystems Identification Manual No. 6. United States Environmental Protection Agency Project # 18050, Contract # 14-12-894, April 1972. x + 82 pp.
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Page edits:

    August 08, 2016
    • Added new records of Plateumaris (Chrysomelidae, Donaciinae) from publication records in Askevold 1991
    August 04, 2016
    • Added new records of Chrysomelidae from publication records in Kölsch & Pedersen 2008
    January 16, 2013
    • Added Phaenonotum exstriatum to list; Smetana (1988) indicates a semi-aquatic ecology, found along water edges; Willston (1970) found it Sphagnum bog mat
    • Added species of Cercyon that Smetana (1988) considers associated with aquatic habitats (peri- to semi-aquatic)
    November 21, 2012
    • Beginning to keep track of changes to the web site pages
    • Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page.

Page created: November 01, 2002; Last update May 12, 2017 (EB)