Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

Plecoptera (Stoneflies) of Michigan

Key to the Plecoptera of Michigan

Introduction

This list provides a recent update to the Plecoptera of Michigan, and is based mainly on Grubbs & Bright (2001) and a more recent update (Grubbs et al. 2012). There are currently about 32 genera and about 71 species recorded for Michigan, with at least an additional 12 species likely to be found based on existing records from nearby state and provincial collection localities. Future sample processing and more thorough sampling across the state - particularly across the Upper Peninsula - will undoubtedly increase the taxonomic representation of Plecoptera in Michigan. An species list for North America is maintained by Dr. Bill Stark at North American Plecoptera has been superceded by the Plecoptera Species File, which has world-wide coverage and many links to older literature.

Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

CAPNIIDAE Banks, 1900 - Snowflies

Allocapnia Claassen, 1928: 667
Allocapnia granulata (Claassen, 1924: 44 as Capnella)
Allocapnia illinoensis Frison, 1935: 365 - IL, IN, MN, OH, ON, WI (expect in the western UP near WI border)
Allocapnia minima (Barnston, 1848: 388 as Perla.)
(syn.) Capnella incisura Claassen, 1924: 45
(syn.) Allocapnia incisura (Claassen, 1924)
Allocapnia nivicola (Fitch, 1847: 274-284 as Perla) - IL, IN, OH, WI
Allocapnia pygmaea (Burmeister, 1839: 874 as Semblis

(syn.) Allocapnia torontonensis Ricker, 1935: 257
Allocapnia recta (Claassen, 1924: 44 as Capnella) - IL, IN, OH, ON, WI
Allocapnia vivipara (Claassen, 1924: 46 as Capnella)

Capnia Pictet, 1841: 320
Capnia vernalis Newport, 1848: 388

(syn.) Perla vernalis Barnston, in Newport 1848
(syn.) Nemoura tenuis Walker, 1852: 182 (not tenuis of Pictet)
(syn.) Capnia limata Frison, 1944: 155

Capnura Banks, 1900: 245
Capnura manitoba (Claassen, 1924: 54 as Capnia) - Montmorency Co., Hunt Creek, 44.8640°N, - 84.1555°W, April 4, 1939, J.W. Leonard, ♂, 2 nymphs (INHS)

Paracapnia Hanson, 1946: 193-249
Paracapnia angulata Hanson, 1961: 25-30 - Yanoviak and McCafferty 1996: 204
Paracapnia opis (Newman, 1839: 32-37,84-90 as Chloroperia)
(syn.) Paracapnia curvata Hanson, 1946: 237

CHLOROPERLIDAE Okamoto, 1912 - Sallflies

Alloperla Banks, 1906: 175
Alloperla atlantica Baumann, 1974: 260
Alloperla banksi Frison, 1942: 343
Alloperla leonarda Ricker, 1952: 176


Haploperla Navas, 1934: 1-16
Haploperla brevis (Banks, 1895: 314 as Chloroperla) - Yanoviak and McCafferty 1996: 204
(comb.) Alloperla brevis (Banks, 1907)
(nom. nud.) Chloroperla cydippe Newman, 1839: 88
(comb.) Hastaperla cydippe (Needham and Claassen, 1925)
(syn.) Hastaperla calcarea Ricker, 1935: 200
(syn.) Isopteryx oculata Klapálek, 1923: 29
Haploperla orpha (Frison, 1937: 91 as Chloroperla)
(comb.) Hastaperla orpha (Frison, 1937) various authors

Suwallia Ricker, 1943: 139
Suwallia marginata (Banks, 1897: 21 as Chloroperla) - see Grubbs et al. 2012

LEUCTRIDAE Klapálek, 1905 - Needleflies

Leuctra Stephens, 1836: 144
Leuctra ferruginea (Walker, 1852: 183 as Nemoura)

(syn.) Leuctra decepta Claassen, 1923: 260
Leuctra rickeri James, 1976: 882-884 - Grubbs et al. 2012: 168
Leuctra sibleyi Claassen, 1923: 262 - IL, IN, OH, ON, WI
Leuctra tenella Provancher, 1878: 124-128 - Yanoviak and McCafferty 1996: 204

(syn.) Leuctra hamula Claassen, 1923: 261
(syn.) Leuctra tenuis Provancher, 1876: 218

NEMOURIDAE Newman, 1853 - Forest stoneflies

Amphinemura Ris, 1902: 384 (subfamily Amphinemurinae Baumann, 1975: 11)
Amphinemura delosa (Ricker, 1952: 18 as Nemoura (Amphinemura)
Amphinemura palmeni (Koponen, 1917: 13 as Nemoura) - Yanoviak and McCafferty 1996: 204 (as A. linda); Boumans & Baumann 2012
(syn.) Nemoura (Amphinemura) linda Ricker, 1952: 22

Nemoura Latreille, 1796: 101
Nemoura trispinosa Claassen, 1923: 289

Ostrocerca Ricker, 1952: 38
Ostrocerca albidipennis (Walker, 1852: 191 as Nemoura)
(syn.) Nemoura serrata Claassen, 1923: 289

Paranemoura Needham and Claassen, 1925: 288
Paranemoura perfecta (Walker, 1852: 191 as Nemoura) - Grubbs et al. 2012: 165
(syn.) Nemoura punctipennis Claassen, 1923: 291

Prostoia Ricker, 1952: 47
Prostoia completa (Walker, 1852: 191 as Nemoura) - Yanoviak and McCafferty 1996: 204
(syn.) Nemoura glabra Claassen, 1923: 281
Prostoia similis (Hagen, 1861: 34 as Taeniopteryx)
(syn.) Nemoura divergens Claassen, 1923: 282

Shipsa Ricker, 1952: 49
Shipsa rotunda (Claassen, 1923: 290 as Nemoura)

Soyedina Ricker, 1952: 50
Soyedina vallicularia (Wu, 1923: 58 as Nemoura)

PERLIDAE Latreille, 1802 - Summer stoneflies

Acroneuria Pictet, 1841: 144 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
Acroneuria abnormis (Newman, 1838: 177 as Perla.) - Stark and Gaufin 1976: 228
(syn.) Perla sonans Newport, 1851: 499
(syn.) Acroneuria depressa Needham & Claassen, 1922: 253
(syn.) Acroneuria eidmanni Šámal, 1933: 96
Acroneuria frisoni Stark & Brown, 1991
Acroneuria internata (Walker, 1852: 152 as Perla.)
Acroneuria lycorias (Newman, 1839: 35 as Perla) - Yanoviak and McCafferty 1996: 204
(syn.) Perla navalis Provancher, 1876: 212
(syn.) Perla riparia Provancher, 1876: 213
(syn.) Perla excavata Banks, 1908: 256
(syn.) Perla perbranchiata Neave, 1933: 237
(syn.) Acroneuria perbranchiata (Neave, 1933)

Agnetina Klapálek, 1907: 16 (Perlinae Latreille, 1802; Perlini Latreille, 1802)
Agnetina capitata (Pictet, 1841: 214 as Perla)
(comb.) Phasganophora capitata Illies, 1966
(syn.) Harrisiola abbreviata Banks, 1948: 122
(syn.) Perla americana Banks, 1900: 243
(syn.) Perla hieroglyphica Provancher, 1876: 211
(syn.) Perla illustris Banks, 1908: 256
(syn.) Perla innota Banks, 1918: 6
(syn.) Perla marginipes Provancher, 1876: 212
(syn.) Harrisiola modesta Banks, 1948: 121
(syn.) Harrisiola nigrescens Banks, 1948: 119
(syn.) Perla tristis Hagen, 1861: 22
Agnetina flavescens (Walsh, 1862: 363 as Perla) - Grubbs et al. 2012: 166
(syn.) Harrisiola klapaleki Banks, 1948: 121

Attaneuria Ricker, 1954: 39 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
Attaneuria ruralis (Hagen, 1861: 18 as Perla (Acroneuria)) - Grubbs et al. 2012: 167 (may be locally extinct from Augusta Creek, site of Fischer's record of Gull Lake Biological Station, 42.3633N, -85.3535N, from July 3, 1957; extirpiated from IL, IN and OH; also recorded from ON and WI)
(comb.) Acroneuria ruralis Claassen, 1940
(syn.) Perla attenuata Banks, 1905: 55
(syn.) Perla quebecensis Provancher, 1876: 211
(syn.) Perla rupinsulensis Walsh, 1862 : 363

Neoperla Needham, 1905: 108 (Perlinae Latreille, 1802; Neoperlini Enderlein, 1909)
Neoperla catharae Stark & Baumann, 1978: 98 - Grubbs et al. 2012: 166
Neoperla occipitalis (Pictet, 1841: 254 as Perla) - IL, IN, OH, ON, WI
(syn.) Neoperla freytagi Stark & Baumann, 1978: 101
Neoperla mainensis Banks, 1948: 124 - IL, IN, ON
Neoperla stewarti Stark & Baumann, 1978: 104 - Grubbs et al. 2012: 166


Paragnetina Klapálek, 1907: 17 (Perlinae Latreille, 1802; Perlini Latreille, 1802)
Paragnetina media (Walker, 1852: 145 as Perla) - Yanoviak and McCafferty 1996: 204, common in collections
(syn.) Acroneuria salvenlini Ricker, 1935: 261
(comb.) Togoperla media Claassen, 1940

Perlesta Banks, 1906: 222 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
Perlesta decipiens (Walsh, 1862: 364 as Perla) - Needham 1908: 262 (as Chloroperla brunneipennis); Grubbs et al. 2012: 166

(syn.) Perla (Chloroperla) brunnipennis Walsh, 1862: 367
(syn.) Perlesta costalis Klapálek, 1921: 150
(syn.) Isoperla texana Banks, 1914: 611
Perlesta ephelida Grubbs & DeWalt, 2012: 4 - Grubbs and Dewalt 2012: 4
Perlesta golconda DeWalt & Stark, 1998: 315 - Grubbs and Dewalt 2012: 166
Perlesta lagoi Stark, 1989: 283 - Grubbs and Dewalt 2012: 166
Perlesta ouabache Grubbs & DeWalt, 2011: 75 - IL, IN, OH (Grubbs & Dewalt 2011)
Perlesta xube Stark & Rhodes, 1997: 92 - IL
Perlesta WI-1 Grubbs & DeWalt, 2012: 170, to be named after female and eggs obtained

Perlinella Banks, 1900: 242 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
Perlinella drymo (Newman, 1839: 86 as Isogenus) - Bright 1996; Grubbs & Bright 2001; Grubbs et al. 2012
(syn.) Perla elongata Walsh, 1862: 366
(syn.) Perla trivittata Banks, 1895: 313
Perlinella ephyre (Newman, 1839: 87 as Chloroperla) - Grubbs & Bright 2001; Grubbs et al. 2012
(comb.) Atoperla ephyre Claassen, 1931
(syn.) Perla producta Walsh, 1862: 365
(syn.) Perla fumipennis Walsh, 1862: 366
(syn.) Atoperla consors Banks, 1948: 123

PERLODIDAE Klapálek, 1909 - Spring stoneflies

Arcynopteryx Klapálek, 1904: 7 (Subfamily Perlodinae)
Arcynopteryx dichroa (McLachlan, 1872: 52 as Dictyopteryx) - Grubbs et al. 2012 (166, see notes on p. 168 regarding previous listing as Arcynopteryx compacta in Grubbs & Bright 2001)
(syn.) Perlodes anisoptera Navás, 1923: 35
(comb.) Arcynopteryx dichroa (Navás, 1923) Illies, 1966
(syn.) Perlodes arnaizi Navás, 1914: 29
(comb.) Arcynopteryx arnaizi (Navás, 1914) Illies, 1966
(syn.) Arcynopteryx brachifer Bengtsson, 1933: 21
(syn.) Arcynopteryx carpathica Klapálek, 1906: 155
(syn.) Arcynopteryx compacta Klapálek, 1904: 7
(syn.) Dictyopteryx dovrensis Morton, 1901: 146
(comb.) Arcynopteryx dovrensis (Klapálek, 1904) Illies, 1966
(syn.) Arcynopteryx ignota Smith, 1917: 479
(syn.) Arcynopteryx inornata Smith, 1917: 480
(syn.) Perlodes lineata Smith, 1917: 476
(comb.) Arcynopteryx lineata (Smith, 1917) Illies, 1966
(syn.) Perlodes margarita Alexander, 1936: 24
(comb.) Arcynopteryx margarita (Alexander, 1936) Illies, 1966
(syn.) Arcynopteryx minor Klapálek, 1912: 22
(syn.) Dictyopteryx norwegica Kempny, 1900: 86
(comb.) Arcynopteryx compacta (Kempny, 1900) Klapálek, 1904
(syn.) Arcynopteryx pyrenaica Despax, 1927: 491
(syn.) Arcynopteryx ringdahli Bengtsson, 1933: 18
(syn.) Perlodes slossonae Banks, 1914: 608
(comb.) Arcynopteryx slossonae (Banks, 1914) Illies, 1966
(syn.) Dictyopteryx transsylvanica Klapálek, 1901: 9
(comb.) Arcynopteryx transsylvanica (Klapálek, 1901) Klapálek, 1904

Clioperla Needham and Claassen, 1925 (Subfamily Isoperlinae)
Clioperla clio (Newman, 1839: 86 as Isogenus)
(comb.) Isoperla clio Frison, 1935: 439
(syn.) Isoperla confusa Frison, 1935: 441

Cultus Ricker 1952: 95 (Perlodinae Klapálek, 1909; Diploperlini Stark & Szczytko, 1984)
Cultus decisus decisus (Walker, 1852: 170 as Perla (Chloroperla) decisa)
(comb.) Perla (Perla) decisa Hagen, 1861
(comb.) Isogenus (Cultus) decisus Ricker, 1952

Helopicus Ricker, 1952: 100 (Perlodinae Klapálek, 1909; Perlodini Klapálek, 1909)
Helopicus nalatus (Frison, 1942: 293 as Hydroperla) - Sandberg & Stewart 2003: 74 (Little Muskegon River, 4.8 Km E of Morley, Mecosta Co., Michigan, 09- IV-1999); note Grubbs et al. 2010: 167 presumes Huron River populations are extinct
(comb.) Isogenus (Helopicus) nalatus Ricker, 1952: 100

Hydroperla Frison, 1935: 417 (Perlodinae Klapálek, 1909; Perlodini Klapálek, 1909)
Hydroperla crosbyi (Needham and Claassen, 1925: 79 as Perla)

Isogenoides Klapálek, 1912: 57 (Perlodinae Klapálek, 1909; Perlodini Klapálek, 1909)
Isogenoides doratus (Frison, 1942: 295 as Hydroperla) - Yanoviak and McCafferty 1996: 204
(syn.) Isogenus (Isogenoides) krumholzi Ricker, 1952: 112
Isogenoides frontalis (Newman, 1838: 178 as Isogenus) - Yanoviak and McCafferty 1996: 204
(syn.) Isogenoides hudsonicus Hanson, 1943: 662
(comb.) Isogenus frontalis Needham and Claassen, 1925 (in part)
Isogenoides olivaceus (Walker, 1852: 144 as Perla) - Yanoviak and McCafferty 1996: 204
(syn.) Perla sulcata Provancher, 1876: 213
(comb.) Perla olivacea Claassen, 1940
(comb.) Isogenus olivacea Ricker, 1938
(comb.) Hydroperla olivacea Frison, 1942

Isoperla Banks, 1906: 175 (Isoperlinae Frison, 1942: 311)
Isoperla bilineata (Say, 1823: 165 as Sialis)
Isoperla cotta Ricker, 1952: 144 - Yanoviak and McCafferty 1996:204
Isoperla decepta Frison, 1935: 447 - Szcytko & Kondratrieff 2015:80
Isoperla dicala Frison, 1942: 321
Isoperla frisoni Illies, 1966: 402 - Yanoviak and McCafferty 1996:204
Isoperla lata Frison, 1942: 334 - Yanoviak and McCafferty 1996:204
Isoperla marlynia Needham and Claassen, 1925: 148
Isoperla montana (Banks, 1898: 199 as Chloroperla) - IN, MN, OH, ON
Isoperla nana (Walsh, 1862: 367 as Chloroperla)
Isoperla orata Frison, 1942: 323 - MN, OH
Isoperla richardsoni Frison, 1935: 459 - Yanoviak and McCafferty 1996:204; Grubbs et al. 2012:166
Isoperla signata (Banks, 1902: 124 as Perlinella)
(syn.) Pictetia bimaculata Banks, 1948: 122
(comb.) Walshiola signata Banks, 1948
Isoperla slossonae (Banks, 1911: 335 as Perla)
(syn.) Clioperla annecta Needham and Claassen, 1925: 140
Isoperla transmarina (Newman, 1838: 499 as Chloroperla) - Frison 1942, Yanoviak and McCafferty 1996:204
(syn.) Isoperla ventralis Banks, 1908: 66
(syn.) Isoperla fumosa Neave, 1933: 235

PTERONARCYIDAE Newman, 1853 - Salmonflies

Pteronarcys Newman, 1838: 175

Pteronarcys dorsata (Say, 1823: 164 as Sialis) - Bright coll.; Yanoviak and McCafferty 1996: 204; widespread in state; see also Nelson 2000: 32-33

(syn.) Pteronarcys flavicornis Provancher, 1876: 191
(syn.) Pteronarcys frigida Gerstaecker, 1873: 65
(syn.) Pteronarcys insignis (Pictet, 1841: 123 as Kollaria)
(syn.) Pteronarcys labradoriensis Šámal, 1933: 95
(syn.) Pteronarcys nobilis Hagen, 1861: 15; (Frison 1935 INHSB 20: 336 has it as P. pictetii)
(syn.) Pteronarcys rectus Provancher, 1876: 189
(syn.) Pteronarcys regalis Newman, 1838: 176
(syn.) Pteronarcys shelfordi Frison, 1934: 25
Pteronarcys pictetii Hagen, 1873: 286
(misid.) Pteronarcys nobilis: Smith 1917, Needham and Claassen 1925, Frison 1935

TAENIOPTERYGIDAE Klapálek, 1905 - Taeniopterygid winter stoneflies

Oemopteryx Klapálek, 1902: 179 (subfamily Brachypterainae Zwick, 1973)
Oemopteryx glacialis (Barnston, in Newport, 1849: 389 as Nemoura)
(syn.) Taeniopteryx (Oemopteryx) alex Hanson, 1938: 79
(syn.) Perla chicoutimiensis Provancher, 1878: 75

Strophopteryx Frison, 1929: 374 (subfamily Brachypterainae Zwick, 1973)
Strophopteryx fasciata (Burmeister, 1839: 875 as Semlis)
(syn.) Taeniopteryx frigida Hagen, 1861: 35
(comb.) Tachiopteryx (Strophopteryx) fasciata Frison, 1942
(comb.) Brachyptera fasciata Harden & Mickel, 1952

Taeniopteryx Pictet, 1841: 345 (subfamily Taeniopteryginae Klapálek, 1905: 27)
Taeniopteryx burksi Ricker & Ross, 1968: 1425 - Yanoviak and McCafferty 1996: 204
Taeniopteryx nivalis Fitch, 1847: 279
Taeniopteryx parvula Banks, 1918: 7

References - Distribution

Alexander KD, Stewart KW. 1999. Revision of the genus Suwallia Ricker (Plecoptera: Chloroperlidae). Transactions of the American Entomological Society 125(3):185-250.
Banks N. 1895. New neuropteroid insects. Transactions of the American Entomological Society 22: 313-316.
Banks N. 1897. New North American neuropteroid insects. Transactions of the American Entomological Society 24: 21-31.
Banks N. 1898. Descriptions of new North American neuropteroid insects. Transactions of the American Entomological Society 25: 199-200.
Banks N. 1900. New genera and species of Nearctic Neuropteroid insects. Transactions of the American Entomological Society 26: 239-259.
Banks N. 1908. Neuropteroid insects, notes and descriptions. Transactions of the American Entomological Society 34:255-267.
Banks N. 1948. Notes on Perlidae. Psyche 55: 113-130.
Baumann RW. 1974. What is Alloperla imbecilla (Say)? Designation of a neotype, and a new Alloperla from eastern North America (Plecoptera: Chloroperlidae). Proceedings of the Biological Society of Washington 87(24):257-264.
Boumans L, Baumann RW. 2012. Amphinemura palmeni is a valid Holarctic stonefly species (Plecoptera: Nemouridae). Zootaxa 3537: 59-75.
Claassen PW. 1924. New species of North American Capniidae (Plecoptera). Canadian Entomologist 56: 43-48, 54-57.
Claassen PW. 1940. A catalogue of the Plecoptera of the world. Memoirs of the Cornell University Agricultural Experimental Station 232:1-235.
Frison TH. 1935. The stoneflies, or Plecoptera, of Illinois. Bulletin of the Illinois Natural History Survey 20:281-471.
Frison TH. 1937. Descriptions of Plecoptera with special reference to the Illinois species. Bulletin of the Illinois Natural History Survey 21:78-99.
Frison TH. 1942. Studies of North American Plecoptera, with special reference to the fauna of Illinois. Bulletin of the Illinois Natural History Survey 22:235-355.
Grubbs SA, Bright E. 2001. Arcynopteryx compacta (Plecoptera: Perlodidae), a Holarctic stonefly confirmed from Lake Superior, with a review and first checklist of the stoneflies of Michigan. The Great Lakes Entomologist 34(2):77-84.
Grubbs SA, Pessino M, DeWalt RE. 2012. Michigan Plecoptera (Stoneflies): distribution patterns and an updated species list. Illiesia 8(18):162-173.
Grubbs SA, DeWalt RE. 2011. Perlesta ouabache, a new species of stonefly (Plecoptera: Perlidae) from Indiana, USA. Aquatic Insects 33(1): 75-79.
Grubbs SA, DeWalt RE. 2012. Perlesta ephelida, a new Nearctic stonefly species (Plecoptera, Perlidae). Zookeys 194: 1-15.
Hanson JF. 1943. Records and descriptions of North American Plecoptera. Part III. Notes on Isogenoides. American Midland Naturalist 29: 657–669.
Hitchcock SW. 1974. Guide to the Insects of Connecticut. Part VII. The Plecoptera or stoneflies of Connecticut. State Geological and Natural History Survey of Connecticut Bulletin 107:1-262.
Needham JG, Claassen PW. 1925. A Monograph of the Plecoptera or Stoneflies of America North of Mexico. Entomological Society of America, The Thomas Say Foundation, Lafayette, Indiana: 1-397.
Nelson CH. 2000. Pteronarcyidae (the Salmonflies), pp. 29-39 in Stark BP, Armitage BJ (eds.), Stoneflies (Plecoptera) of Eastern North America, Volume 1: Pteronarcyidae, Peltoperlidae, and Taeniopterygidae. Ohio Biological Survey Bulletin New Series Volume 14, Number 1. vii + 100p.
Sandberg JB, Stewart KW. 2003. Continued studies of drumming in North American Plecoptera; Evolutionary implications, pp. 73-81 in E. Gaino (ed.). Research update on Ephemeroptera and Plecoptera. University of Perugia, Perugia, Italy.
Sandberg JB, Stewart KW. 2005. Holomorphology and Systematics of the Stonefly Genus Isogenoides (Plecoptera: Perlodidae). Transactions of the American Entomological Society 131(3+4): 269-345.
Stewart KW. 2000. Taeniopterygidae (the Willowflies), pp. 55-88 in Stark BP, Armitage BJ (eds.), Stoneflies (Plecoptera) of Eastern North America, Volume 1: Pteronarcyidae, Peltoperlidae, and Taeniopterygidae. Ohio Biological Survey Bulletin New Series Volume 14, Number 1. vii + 100p.
Stewart KW, Stark BP. 2002. Nymphs of North American stonefly genera. Second Edition. The Caddis Press: Columbus, Ohio. xii + 510
Szczytko SW, Kondratieff BC. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia 1:1-289.
Surdick RF. 1985. Nearctic genera of Chloroperlidae (Plecoptera: Chloroperlidae). Illinois Biological Monographs 54:1-146.
Teslenko VA. 2012. A taxonomic revision of the genus Arcynopteryx Klapálek, 1904 (Plecoptera, Perlodidae). Zootaxa 3329:1-18.
USGS Northern Prairie Wildlife Rearch Center - web site managed by B. Kondratieff (accessed August 6, 2006)
Yanoviak SP, McCafferty WP. 1996. Comparison of macroinvertebrate assemblages inhabiting pristine streams in the Huron Mountains of Michigan, USA. Hydrobiologia 330: 195-211.

References - Taxonomy

Alexander CP. 1936. A new species of Perlodes from the White Mountains, New Hampshire (family Perlidae; order Plecoptera). Bulletin of the Brooklyn Entomological Society 31: 24-27.
Alexander KD, Stewart KW. 1999. Revision of the genus Suwallia Ricker (Plecoptera: Chloroperlidae). Transactions of the American Entomological Society 125(3):185-250.
Banks N. 1895. New neuropteroid insects. Transactions of the American Entomological Society 22: 313-316.
Banks N. 1897. New North American neuropteroid insects. Transactions of the American Entomological Society 24: 21-31.
Banks N. 1898. Descriptions of new North American neuropteroid insects. Transactions of the American Entomological Society 25: 199-200.
Banks N. 1900. New genera and species of Nearctic neuropteroid insects. Transactions of the American Entomological Society 26: 239-259.
Banks N. 1902. Notes and descriptions of Perlidae. The Canadian Entomologist 34: 123-125.
Banks N. 1905. New genera and species of Perlidae. Psyche 12: 55-57.
Banks N. 1906. On the perlid genus Chloroperla. Entomological News 17: 174-175.
Banks N. 1907. Catalogue of the neuropteroid insects (except Odonata) of the United States. American Entomological Society, 53 pp.
Banks N. 1908. Neuropteroid insects, notes and descriptions. Transactions of the American Entomological Society 34: 255-267.
Banks N. 1911. Descriptions of new species of North American neuropteroid insects. Transactions of the American Entomological Society 37: 335-360.
Banks N. 1914. New neuropteroid insects, native and exotic. Proceedings of the Academy of Natural Science, Philadelphia 66: 608-632.
Banks N. 1918. New neuropteroid insects. Bulletin of the Museum of Comparative Zoology 62: 3-22.
Banks N. 1948. Notes on Perlidae. Psyche 55: 113-130.
Barnston. 1848. In Newport G. Postscript to On the Anatomy and affinities of Pteronarcys regalis, Newm. Proceedings of the Linnean Society of London 36: 388.
Baumann RW. 1974. What is Alloperla imbecilla (Say)? Designation of a neotype, and a new Alloperla from eastern North America (Plecoptera: Chloroperlidae). Proceedings of the Biological Society of Washington 87(24):257-264.
Bengtsson S. 1933. Plecopterologische Studien. Lunds Universität Arsskrift (2)29: 1-50
Boumans L, Baumann RW. 2012. Amphinemura palmeni is a valid Holarctic stonefly species (Plecoptera: Nemouridae). Zootaxa 3537: 59-75.
Burmeister H. 1839. Handbuch der Entomologie. Vol 2. Enslin: Berlin, pp. 397-1050.
Claassen PW. 1923. New species of North American Plecoptera. The Canadian Entomologist 55(11): 257-263.
Claassen PW. 1924. New species of North American Capniidae (Plecoptera). The Canadian Entomologist 56(1): 43-48, 54-57.
Claassen PW. 1928. Additions and corrections to the monograph on the Plecoptera of North America. Annals of the Entomological Society of America 21(4) : 667-668.
Claassen PW. 1940. A catalogue of the Plecoptera of the world. Memoirs of the Cornell University Agricultural Experimental Station 232:1-235.
Despax R. 1927. Arcynopteryx pyrenaica, Plécoptètr nouveau des Pyrénées françaises, Bulletin de la Société d'histoire Naturelle de Toulouse 56:491-495.
DeWalt RE, Stark BP, Harris MA. 1998. Perlesta golconda (Plecoptera: Perlidae), a new stonefly species from Illinois. Entomological News 109: 315-317.
Fitch A. 1847. Winter insects of eastern New York. American Journal of Agricultural Science 5: 274-284.
Frison TH. 1929. Fall and winter stoneflies or Plecoptera of Illinois. Bulletin of the Illinois Natural History Survey 18: 345-409.
Frison TH. 1934. Four new species of stoneflies from North America (Plecoptera). The Canadian Entomologist 66: 25-30.
Frison TH. 1935. The stoneflies, or Plecoptera, of Illinois. Bulletin of the Illinois Natural History Survey 20:281-471.
Frison TH. 1937. Descriptions of Plecoptera with special reference to the Illinois species. Bulletin of the Illinois Natural History Survey 21:78-99.
Frison TH. 1942. Studies of North American Plecoptera, with special reference to the fauna of Illinois. Bulletin of the Illinois Natural History Survey 22:235-355.
Gerstaecker A. 1873. Über Pteronarcys Newm. und eine zweite, im Imago-Stadium mit Tracheen-kiemen versehene Perlarien-Gattung, in Zur Morphologie der Orthoptera amphibiotica. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1873: 60-74.
Grubbs SA, DeWalt RE. 2011. Perlesta ouabache, a new species of stonefly (Plecoptera: Perlidae) from Indiana, USA. Aquatic Insects 33(1): 75-79.
Grubbs SA, DeWalt RE. 2012. Perlesta ephelida, a new Nearctic stonefly species (Plecoptera, Perlidae). Zookeys 194: 1-15.
Harden PH, Mickel CE. 1952. The stoneflies of Minnestoa (Plecoptera). Technical Bulletin of the University of Minnesota Agricultural Experimental Station 201: 1-84.
Hanson JF. 1938. Studies on the Plecoptera of North America. I. Bulletin of the Brooklyn Entomological Society 33: 79-83.
Hanson JF. 1943. Records and descriptions of North American Plecoptera. Part III. Notes on Isogenoides. American Midland Naturalist 29: 657–669.
Hanson JF. 1946. Comparative morphology and taxonomy of the Capniidae (Plecoptera). American Midland Naturalist 35(1):193-249
Hanson JF. 1961. Studies on the Plecoptera of North America: VIII. The identity of the species of Paracapnia. Bulletin of the Brooklyn Entomological Society 56(2): 25-30.
Hagen HA. 1861. Synopsis of the Neuroptera of North America, with a list of the South American species. Smithsonian Miscellaneous Collections 4:1-347.
Hagen HA. 1873. Perlina. In Report on the Pseudoneuroptera and Neuroptera of North America in the collection of the late Th. W. Harris. Proceedings of the Boston Society of Natural History, Boston 15: 281-290.
Hitchcock SW. 1974. Guide to the Insects of Connecticut. Part VII. The Plecoptera or stoneflies of Connecticut. State Geological and Natural History Survey of Connecticut Bulletin 107:1-262.
Illies J. 1966. Katalog der rezenten Plecoptera. Das Tierreich 82. Walter de Gruyter, Berlin. 631 pp.
James AM. 1976. Two new species of Leuctra, with notes on the ferruginea group (Plecoptera: Leuctridae). Annals of the Entomological Society of America 69: 882-884.
Klapálek F. 1901. Über neue und wenig bekannte Arten der paläarktischen Neuropteroiden., Bulletin of the International Academy of Sciences, Bohéme (Sci. math.-nat.), Praha 7: 1-14.
Klapálek F. 1902. Zur Kenntniss der Neuropteroiden von Ungarn, Bosnien und Herzegovina. Természetrajzi Füzetek 25: 161–180.
Klapálek F. 1904. Evropske druhy čeledĕ Dictyopterygidae. Rozpravy Česke Akademie (Praha) 2: 1−10.
Klapálek F. 1905. Conspectus Plecopterorum Bohemiae. Casopis Ceskoslovenské Spolecnosti Entomologické 2:27-32.
Klapálek F. 1906. Revision und Synopsis der europäischen Dictyopterygiden. Bulletin of the International Academy of Sciences, Bohéme (Sci. math.-nat.), Praha 11:137-166.
Klapálek F. 1907. Evropske druhy rodu Perla Geoffr. Rozpravy České akademie věd a umění: (Mathematicko-přírodovědecká), Trida II, 16: 1-25.
Klapálek F. 1909. Revise rodu Acronueria Pict. Rozpravy Česke Akademie (Praha) 18: 1−21.
Klapálek F. 1912. Plécoptères I. Fam. Perlodidae. Collections Zoologiques du Baron Edmond de Sélys-Longchamps. Bruxelles 4: 1–66.
Klapálek F. 1914. Analyticka tabulka fam. Perlidae a jeji dvou subfam., Perlinae a Acroneuriinae (Plecoptera). Casopis Ceskoslovenské Spolecnosti Entomologické 2:53-69.
Klapálek F. 1923. Plécoptères nouveaux. Annales de la Société entomologique de Belgique 63: 21-29.
(syn.) Dictyopteryx compacta Kempny, 1900
Koponen J. 1917. Plecopterologische Studien I: Die Plecopteren-Arten Finlands. Acta Societatis pro fauna et flora Fennica 44: 1–18.
Klapálek F. 1921. Plécoptères nouveaux. Ann. Annales de la Société entomologique de Belgique (Bruxelles), 61:57-67, 146-150, 320, 327.
Kempny P. 1900. Beitrag zur Perliden- und Trichopteren-Fauna Südtirols. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 50: 254-258.
Latreille PA. 1796. Nevropteres. pp. 96-104 in Prc'cis des Caractires Generiques Insectes, Disposes dans un Ordre Naturel. Bordeaux.
Latreille PA. 1802. Histoire naturelle, générale et particulière des crustacés et des insectes: ouvrage faisant suite aux oeuvres de Leclerc de Buffon, et partie du cours complet d'histoire naturelle rédigé 3. 467 pp.
McLachlan R. 1872. Perlides. In Non-Odonates (Second part, Materiaux pour une fauna nevropterologique de l'Asie septentrionale), by de Selys-Longchamps & McLachlan. Annales de la Société entomologique de Belgique 15: 47−55.
Morton KJ. 1901. Perlidae taken in Norway in June and July, 1900, with remarks on certain Arctic forms. The Entomologist's Monthly Magazine 37:146-148.
Navás RPL. 1923. Plecópteros, in Algunos insectos del Museo de Paris. Revista de la Academia de Ciencias de Zaragoza 7: 33-44.
Navas RPL. 1934. Plecoptera, pp. 9-12 in Névroptères et insectes voisin (Chine et pays environants). Notes d'Entomologie Chinoise 2.
Neave F. 1933. Some new stoneflies from western Canada. The Canadian Entomologist 65: 235-238.
Needham JG. 1905. New genera and species of Perlidae. Proceedings of the Biological Society of Washington 18: 107-110.
Needham JG, Claassen PW. 1922. The North American species of the genus Acroneuria (Order Plecoptera). The Canadian Entomologist 65: 235-238.
Needham JG, Claassen PW. 1925. A Monograph of the Plecoptera or Stoneflies of America North of Mexico. Entomological Society of America, The Thomas Say Foundation, Lafayette, Indiana: 1-397.
Nelson CH. 2000. Pteronarcyidae (the Salmonflies), pp. 29-39 in Stark BP, Armitage BJ (eds.), Stoneflies (Plecoptera) of Eastern North America, Volume 1: Pteronarcyidae, Peltoperlidae, and Taeniopterygidae. Ohio Biological Survey Bulletin New Series Volume 14, Number 1. vii + 100p.
Newman E. 1838. Entomological notes. Entomological Magazine 5: 168-181, 372-402, 483-500.
Newman E. 1839. On the synonymy of Perlites, together with brief charcters of the old, and of a few new species. The Annals and Magazine of Natural History 3: 32-37, 84-90.
Newman E. 1853. Proposed division of Neuroptera into two classes. The Zoologist 11: 82-204.
Newport G. 1848/1849. Postscript to Mr. Newport's paper on Pteronarcys regalis. Proceedings of the Linnean Society of London 1: 387-389.
Newport G. 1851. On the anatomy and affinities of Pteronarcys regalis Newm.: with a postscript, containing descriptions of some American Perlidac, together with notes on their habits. Transactions of the Linnean Society of London 20(3): 447-452.
Okamoto H. 1912. Erste Beitrag zur Kenntnis der Japanischen Plecopteren. Transactions of the Sapporo Natural History Society 4: 105-170.
Pictet FJ. 1841. Histoire naturelle générale et particulière des insects Névroptères. Famille des Perlides. 1. Partie. Kessmann, Geneve. 423 pp.
Provancher L. 1876. Petit faune entomologique du Canada. Les Névroptères. Le Naturalist Canadienne 8: 177-191, 209-218.
Provancher L. 1878. Petit faune entomologique du Canada et Particulierement de la Province de Quebec. Troiseemi ordre, Les Névroptères. Le Naturalist Canadienne 10: 55-162.
Ricker WE. 1935. New Canadian perlids (part II). The Canadian Entomologist 67: 256-264.
Ricker WE. 1938. Notes on specimens of American Plecoptera in European collections. Transactions of the Royal Canadian Institute 22: 129-156.
Ricker WE. 1943. Stoneflies of southwestern British Columbia. Indiana University Publication, Science Series 12: 1-145.
Ricker WE. 1952. Systematic studies of Plecoptera. Indiana University Publication, Science Series 18: 1-200.
Ricker WE. 1954. Nomenclatorial notes on Plecoptera. Proceedings of the Entomological Society of British Columbia 51: 37-39.
Ricker WE, Ross HH. 1968. North American species of Taeniopteryx (Plecoptera, Insecta). Journal of the Fisheries Research Board of Canada 25: 1423-1439.
Ris F. 1902. Die Schweizerischen Arten der Perliden-Gattung Nemura. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 10: 378-384.
Šámal J. 1933. Beiträge zur Kenntnis der Plecopterenfauna Südlabradors. Zoologischer Anzeiger 102(3/4): 95-100.
Sandberg JB, Stewart KW. 2005. Holomorphology and Systematics of the Stonefly Genus Isogenoides (Plecoptera: Perlodidae). Transactions of the American Entomological Society 131(3+4): 269-345.
Say T. 1823. Description of insects belonging to the order Neuroptera Linn. and Latr. collected by the expedition authorized by J. C. Calhoun, Secretary of War, under the command of Major S. H. Long. Godman's Western Quarterly Report 2: 160-165.
Smith LW. 1917. Studies of North American Plecoptera (Pteronarcinae and Perlodini). Transactions of the American Entomological Society 43: 433-489.
Stark BP, Baumann RW. 1978. New species of nearctic Neoperla (Plecoptera: Perlidae), with notes on the genus. Great Basin Naturalist 38(97-114.
Stark BP, Brown LD. 1991. What is Acroneuria evoluta Klapálek (Plecoptera: Perlidae)? Aquatic Insects 13(1): 29-32.
Stark BP, Rhodes HA. 1997. Perlesta xube, a new stonefly species from Nebraska (Plecoptera: Perlidae). Entomological News 108: 92-96.
Stephens JF. 1836. Illustrations of British entomology; or a synopsis of indigenous insects: containing their generic and specific distinctions with an account of their metamorphoses, times of appearance, localities, food and economy, as far as practicable, with coloured figures (from Westwood) of the rare and more interesting species. Baldwin and Cradock, London. 6: 144.
Szczytko SW, Kondratieff BC. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia 1:1-289.
Surdick RF. 1985. Nearctic genera of Chloroperlidae (Plecoptera: Chloroperlidae). Illinois Biological Monographs 54:1-146.
Teslenko VA. 2012. A taxonomic revision of the genus Arcynopteryx Klapálek, 1904 (Plecoptera, Perlodidae). Zootaxa 3329:1-18.
Walker F. 1852. Catalogue of the specimens of neuropterous insects in the collection of the British Museum. Part I. London. 192 pp.
Walsh BD. 1862. List of the Pseudoneuroptera of Illinois contained in the cabinet of the writer, with descriptions of over forty new species, and notes on their structural affinities. Proceedings of the Academy of Natural Science, Philadelphia 13: 361-402.
Walsh BD. 1863-1864. Observations on certain N.A. Neuroptera by H. Hagen, M.D. of Konigsberg, Prussia; translated from the original French MS and publ. by permission of the author with notes and descriptions of about 20 new species of North American Neuroptera. Proceedings of the Academy of Natural Science, Philadelphia 1863: 168-169.
Wu CF. 1923. Morphology, anatomy and ethology of Nemoura. Bulletin of the Lloyd Library of Botany, Natural History, Pharmacy and Materia Medica, Entomological Series 13: 1-81.
Zwick P. 1973. Insecta: Plecoptera, Phylogenetisches System und Katalog. Das Tierreich, 94. Walter de Gruyter, Berlin. 465 pp.

March 06, 2017
• Reorganized reference sections, splitting them into references pertaining to sources documenting the occurence of a species in Michigan, and another for works that reference authorship of species cited.
March 23, 2015
• Isoperla decepta added to the state species list, as listed in a new publication by Szczytko and Kondratieff (2015)
• Leuctra tenuis, previously listed, removed in Grubbs and Bright (2001) as the record from Yanoviak and McCafferty 1996 (p. 204), based on nymphs, proved unsupported, and its currently known distribution (and in the updated Michigan list by Grubbs et al. (2012) make it unlikely to be recorded in Michigan. Therefore, I have removed this species from the list. I have also removed a number of other species (Allocapnia frisoni, Allocapnia rickeri, Taeniopteryx metequi, Perlesta adena), although regionally recorded, have distributions from only unglaciated areas and are not likely to be found in Michigan.
March 20, 2015
• Paranemoura perfecta and Arcynopteryx dichroa, listed in Grubbs et al. (2012), has also been added to the list. This edit supercedes the edit dated November 04, 2012 (see below) for Arcynopteryx compacta, which is now in synonomy with Arcynopteryx dichroa.
• Leuctra tenuis, previously listed, removed in Grubbs and Bright (2001) as the record from Yanoviak and McCafferty 1996 (p. 204), based on nymphs, proved unsupported, and its currently known distribution (and in the updated Michigan list by Grubbs et al. (2012) make it unlikely to be recorded in Michigan. Therefore, I have removed this species from the list.
January 04, 2013
• Taeniopteryx maura, recorded as new to Michigan by Grubbs and Bright (2001), was based on a prior misidentification of T. burksi, and is removed from the list.
• Isogenoides varians, previously recorded in Michigan by Needham and Claassen (1925: 84) and Hanson (1943), are unsubstantiated without location of specimens or misidentifications; based on it's known distribution, it is unlikely to be found in Michigan (Sandberg and Stewart 2005). It is removed from the list.
December 21, 2012
• Boumans & Baumann (2012) recently synonomyzed Amphinemura linda with Amphinemura palmeni based on morphological and genetic evidence
• Grubbs & DeWalt (2012) described Perlesta ephelida, found in Michigan; previous records of Perlesta shubuta from Michigan (Grubbs and Bright 2001) are of P. ephelida
November 04, 2012
• Beginning to keep track of changes to the web site pages
• Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page
• Updated synonomies per Plecoptera Home Page
• Added Perlesta ouabache (Grubbs & DeWalt, 2011).
• Arcynopteryx compacta (McLachlan, 1872) moved to Skwala Ricker, 1943 (Teslenko 2012).

Page created: October 29, 2002 - Last updated: March 24, 2015 (EB)