Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Plecoptera (Stoneflies) of Michigan

Key to the Plecoptera of Michigan

Introduction

This list provides a recent update to the Plecoptera of Michigan, and is based mainly on Grubbs & Bright (2001) and a more recent update (Grubbs et al. 2012). There are currently about 32 genera and about 69 species recorded for Michigan, with additional species occurring nearby noted with state localities from which certified records have been published. Future sample processing and more thorough sampling across the state - particularly across the Upper Peninsula - will undoubtedly increase the taxonomic representation of Plecoptera in Michigan. An species list dated October 12, 1998 for North America is maintained by Dr. Bill Stark at North American Plecoptera [note: as of March 22, 2015, the link appears to be broken, and is becoming outdated], with current nomenclature maintained at the Plecoptera Home Page.

Genera are donoted in bold green; species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

CAPNIIDAE Banks, 1900 - Snowflies

    Allocapnia Claassen, 1928
      Allocapnia granulata (Claassen, 1924) Capnella (orig.)
      Allocapnia illinoensis Frison, 1935 - IL, IN, MN, OH, ON, WI (expect in the western UP near WI border)
      Allocapnia minima (Barnston, 1848) Perla (orig.)
        (syn.) Allocapnia incisura (Claassen, 1924)
        (syn.) Capnella incisura Claassen, 1924
      Allocapnia nivicola (Fitch, 1847) Perla (orig.) - IL, IN, OH, WI
      Allocapnia pygmaea (Burmeister, 1839) Semblis
        (syn.) Allocapnia torontonensis Ricker, 1935
      Allocapnia recta (Claassen, 1924) Capnella (orig.) - IL, IN, OH, ON, WI
      Allocapnia vivipara (Claassen, 1924) Capnella (orig.)

    Capnia Pictet, 1841
      Capnia vernalis Newport, 1848
        (syn.) Perla vernalis Barnston, 1848
        (syn.) Nemoura tenuis Walker, 1852 (not tenuis of Pictet)

    Capnura Banks, 1900
      Capnura manitoba (Claassen, 1924) Capnia (orig.) - Montmorency Co., Hunt Creek, 44.8640°N, - 84.1555°W, April 4, 1939, J.W. Leonard, ♂, 2 nymphs (INHS);

    Paracapnia Hanson, 1946
      Paracapnia angulata Hanson, 1961 - Yanoviak and McCafferty 1996: 204
      Paracapnia opis (Newman, 1839) Chloroperia (orig.)
        (syn.) Paracapnia curvata Hanson, 1946

CHLOROPERLIDAE Okamoto, 1912 - Sallflies

    Alloperla Banks, 1906
      Alloperla atlantica Baumann, 1974
      Alloperla banksi Frison, 1942
      Alloperla leonarda Ricker, 1952

    Haploperla Navas, 1934
      Haploperla brevis (Banks, 1895) Chloroperla (orig.) - Yanoviak and McCafferty 1996: 204
        (comb.) Alloperla brevis (Banks, 1907)
        (syn.) Chloroperla cydippe Needham & Claassen, 1925
        (syn.) Hastaperla cydippe (Needham & Claassen, 1925)
        (syn.) Hastaperla calcarea Ricker, 1935
        (syn.) Isopteryx oculata Klapálek, 1923
      Haploperla orpha (Frison, 1937) Chloroperla
        (comb.) Hastaperla orpha (Frison, 1937)

    Suwallia Ricker, 1943
      Suwallia marginata (Banks, 1897) Chloroperla (orig.) - see Grubbs et al. 2012

LEUCTRIDAE Klapálek, 1905 - Needleflies

    Leuctra Stephens, 1835
      Leuctra ferruginea (Walker, 1852) Nemoura (orig.)
        (syn.) Leuctra decepta Claassen, 1923
      Leuctra rickeri James, 1976 - Grubbs et al. 2012: 168
      Leuctra sibleyi Claassen, 1923 - IL, IN, OH, ON, WI
      Leuctra tenella Provancher, 1878 - Yanoviak and McCafferty 1996: 204
        (syn.) Leuctra hamula Claassen, 1923
        (syn.) Leuctra tenuis Provancher, 1876

NEMOURIDAE Newman, 1853 - Forest stoneflies

    Amphinemura Ris, 1902 (subfamily Amphinemurinae Baumann, 1975)
      Amphinemura delosa (Ricker, 1952) Nemoura (Amphinemura) (orig.)
      Amphinemura palmeni (Koponen, 1917) Nemoura (orig.) - Yanoviak and McCafferty 1996: 204 (as A. linda); Boumans & Baumann 2012
        (syn.) Nemoura linda Ricker, 1952
        (syn.) Amphinemura linda (Ricker, 1952)

    Nemoura Latreille, 1796
      Nemoura trispinosa Claassen, 1923

    Ostrocerca Ricker, 1952
      Ostrocerca albidipennis (Walker, 1852) Nemoura (orig.)
        (syn.) Nemoura serrata Claassen, 1923

    Paranemoura Needham & Claassen, 1925
      Paranemoura perfecta (Walker, 1852) Nemoura (orig.) - Grubbs et al. 2012: 165
        (syn.) Nemoura punctipennis Claassen, 1923

    Prostoia Ricker, 1952
      Prostoia completa (Walker, 1852) Nemoura (orig.) - Yanoviak and McCafferty 1996: 204
      Prostoia similis (Hagen), 1861 Taeniopteryx (orig.)

    Shipsa Ricker, 1952
      Shipsa rotunda (Claassen, 1923) Nemoura (orig.)

    Soyedina Ricker, 1952
      Soyedina vallicularia (Wu, 1923) Nemoura (orig.)

PERLIDAE Latreille, 1802 - Summer stoneflies

    Acroneuria Pictet, 1841 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
      Acroneuria abnormis (Newman, 1838) Perla (orig.)
        (syn.) Perla sonans Newport, 1851
        (syn.) Acroneuria depressa Needham & Claassen, 1922
        (syn.) Acroneuria eidmanni Šámal, 1933
      Acroneuria frisoni Stark & Brown, 1991
      Acroneuria internata (Walker, 1852) Perla (orig.)
      Acroneuria lycorias (Newman, 1839) Perla (orig.) - Yanoviak and McCafferty 1996: 204
        (syn.) Perla navalis Provancher, 1876
        (syn.) Perla riparia Provancher, 1876
        (syn.) Perla excavata Banks, 1908
        (syn.) Perla perbranchiata Neave, 1933
        (syn.) Acroneuria perbranchiata (Neave, 1933)

    Agnetina Klapálek, 1907 (Perlinae Latreille, 1802; Perlini Latreille, 1802)
      Agnetina capitata (Pictet, 1841) Perla (orig.)
        (comb.) Phasganophora capitata Illies, 1966
        (syn.) Harrisiola abbreviata Banks, 1948
        (syn.) Perla americana Banks, 1900
        (syn.) Perla hieroglyphica Provancher, 1876
        (syn.) Perla illustris Banks, 1908
        (syn.) Perla innota Banks, 1918
        (syn.) Perla marginipes Provancher, 1876
        (syn.) Harrisiola modesta Banks, 1948
        (syn.) Harrisiola nigrescens Banks, 1948
        (syn.) Perla tristis Hagen, 1861
      Agnetina flavescens (Walsh, 1862) Perla (orig.) - Grubbs et al. 2012: 166
        (syn.) Harrisiola klapaleki Banks, 1948

    Attaneuria Ricker, 1954 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
      Attaneuria ruralis (Hagen, 1861) Perla (Acroneuria) (orig.) - Grubbs et al. 2012: 167 (may be locally extinct from Augusta Creek, site of Fischer's record of Gull Lake Biological Station, 42.3633N, -85.3535N, from July 3, 1957; extirpiated from IL, IN and OH; also recorded from ON and WI)
        (comb.) Acroneuria ruralis Claassen, 1940
        (syn.) Perla attenuata Banks, 1905
        (syn.) Perla quebecensis Provancher, 1876
        (syn.) Perla rupinsulensis Walsh, 1862

    Neoperla Needham, 1905(Perlinae Latreille, 1802; Neoperlini Enderlein, 1909)
      Neoperla catharae Stark & Baumann, 1978 - Grubbs et al. 2012: 166
      Neoperla occipitalis (Pictet, 1841) Perla (orig.) - IL, IN, OH, ON, WI
        (syn.) Neoperla freytagi Stark & Baumann, 1978
      Neoperla mainensis Banks, 1948 - IL, IN, ON
      Neoperla stewarti Stark & Baumann, 1978 - Grubbs et al. 2012: 166

    Paragnetina Klapálek, 1907 (Perlinae Latreille, 1802; Perlini Latreille, 1802)
      Paragnetina media (Walker, 1852) Perla (orig.) - Yanoviak and McCafferty 1996: 204, common in collections
        (syn.) Acroneuria salvenlini Ricker, 1935
        (comb.) Togoperla media Claassen, 1940

    Perlesta Banks, 1906 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
      Perlesta decipiens (Walsh, 1862) Perla (orig.) - Needham 1908: 262 (as Chloroperla brunneipennis); Grubbs et al. 2012: 166
        (syn.) Perla (Chloroperla) brunnipennis Walsh, 1862
        (syn.) Perlesta costalis Klapálek, 1921
        (syn.) Isoperla texana Banks, 1914
      Perlesta ephelida Grubbs & DeWalt, 2012 - Grubbs and Dewalt 2012: 166
      Perlesta golconda DeWalt & Stark, 1998 - Grubbs and Dewalt 2012: 166
      Perlesta lagoi Stark, 1989 - Grubbs and Dewalt 2012: 166
      Perlesta ouabache Grubbs & DeWalt, 2011 - IL, IN, OH (Grubbs & Dewalt 2011)
      Perlesta xube Stark & Rhodes, 1997 - IL
      Perlesta WI-1 Grubbs & DeWalt, 2012: 170, to be named after female and eggs obtained

    Perlinella Banks, 1900 (Acroneuriinae Klapálek, 1914; Acroneuriini Klapálek, 1914)
      Perlinella drymo (Newman, 1839) Isogenus (orig.) - Bright 1996; Grubbs & Bright 2001; Grubbs et al. 2012
        (syn.) Perla elongata Walsh, 1862
        (syn.) Perla trivittata Banks, 1895
      Perlinella ephyre (Newman, 1839) Chloroperla (orig.) - Grubbs & Bright 2001; Grubbs et al. 2012
        (comb.) Atoperla ephyre Claassen, 1940
        (syn.) Perla producta Walsh, 1862
        (syn.) Perla fumipennis Walsh, 1862
        (syn.) Atoperla consors Banks, 1948

PERLODIDAE Klapálek, 1909 - Spring stoneflies

    Arcynopteryx Klapálek, 1904 (Subfamily Perlodinae)
      Arcynopteryx dichroa (McLachlan, 1872) Dictyopteryx (orig.) - Grubbs et al. 2012 (166, see notes on 168 regarding previous listing as Arcynopteryx compacta in Grubbs & Bright 2001)
        (syn.) Perlodes anisoptera Navás, 1923
        (comb.) Arcynopteryx dichroa (Navás, 1923) Illies, 1966
        (syn.) Perlodes arnaizi Navás, 1914
        (comb.) Arcynopteryx arnaizi (Navás, 1914) Illies, 1966
        (syn.) Arcynopteryx brachifer Bengtsson, 1933
        (syn.) Arcynopteryx carpathica Klapálek, 1906
        (syn.) Dictyopteryx dovrensis Morton, 1901
        (comb.) Arcynopteryx dovrensis (Klapálek, 1904) Illies, 1966
        (syn.) Arcynopteryx ignota Smith, 1917
        (syn.) Arcynopteryx inornata Smith, 1917
        (syn.) Perlodes lineata Smith, 1917
        (comb.) Arcynopteryx lineata (Smith, 1917) Illies, 1966
        (syn.) Perlodes margarita Alexander, 1936
        (comb.) Arcynopteryx margarita (Alexander, 1936) Illies, 1966
        (syn.) Arcynopteryx minor Klapálek, 1912
        (syn.) Dictyopteryx compacta Kempny, 1900
        (comb.) Arcynopteryx compacta (Kempny, 1900) Klapálek, 1904
        (syn.) Arcynopteryx pyrenaica Despax, 1927
        (syn.) Arcynopteryx ringdahli Bengtsson, 1933
        (syn.) Perlodes slossonae Banks, 1914
        (comb.) Arcynopteryx slossonae (Banks, 1914) Illies, 1966
        (syn.) Dictyopteryx transsylvanica Klapálek, 1901
        (comb.) Arcynopteryx transsylvanica (Klapálek, 1901) Klapálek, 1904

    Clioperla Needham & Claassen, 1925 (Subfamily Isoperlinae)
      Clioperla clio (Newman,1839) Isogenus (orig.)
        (comb.) Isoperla clio Frison, 1935
        (syn.) Isoperla confusa Frison, 1935

    Cultus Ricker 1952 (Perlodinae Klapálek, 1909; Diploperlini Stark & Szczytko, 1984)
      Cultus decisus decisus (Walker, 1852) Perla (orig.)
        (comb.) Perla (Perla) decisa Hagen, 1861
        (comb.) Isogenus (Cultus) decisus Ricker, 1952

    Helopicus Ricker, 1952 (Perlodinae Klapálek, 1909; Perlodini Klapálek, 1909)
      Helopicus nalatus (Frison, 1942) Hydroperla (orig.) - Sandberg & Stewart 2003: 74 (Little Muskegon River, 4.8 Km E of Morley, Mecosta Co., Michigan, 09- IV-1999); note Grubbs et al. 2010: 167 presumes Huron River populations are extinct
        (comb.) Isogenus (Helopicus) nalatus Ricker, 1952

    Hydroperla Frison, 1935 (Perlodinae Klapálek, 1909; Perlodini Klapálek, 1909)
      Hydroperla crosbyi (Needham & Claassen, 1925) Perla (orig.)

    Isogenoides Klapálek, 1912 (Perlodinae Klapálek, 1909; Perlodini Klapálek, 1909)
      Isogenoides doratus (Frison, 1942) Hydroperla - Yanoviak and McCafferty 1996: 204
        (syn.) Isogenus (Isogenoides) krumholzi Ricker, 1952
      Isogenoides frontalis (Newman, 1838) Isogenus (orig.) - Yanoviak and McCafferty 1996: 204
        (comb.) Isogenus frontalis Needham & Claassen, 1925 (in part)
      Isogenoides olivaceus (Walker, 1852) Perla - Yanoviak and McCafferty 1996: 204
        (syn.) Perla sulcata Provancher, 1876
        (comb.) Perla olivacea Claassen, 1940
        (comb.) Isogenus olivacea Ricker, 1938
        (comb.) Hydroperla olivacea Frison, 1942

    Isoperla Banks, 1906 (Isoperlinae Frison, 1942)
      Isoperla bilineata (Say, 1823) Sialis (orig.)
      Isoperla cotta Ricker, 1952 - Yanoviak and McCafferty 1996:204
      Isoperla decepta Frison, 1935 - Szcytko & Kondratrieff 2015:80
      Isoperla dicala Frison, 1942
      Isoperla frisoni Illies, 1966 - Yanoviak and McCafferty 1996:204
      Isoperla lata Frison, 1942 - Yanoviak and McCafferty 1996:204
      Isoperla marlynia Needham & Claassen, 1925
      Isoperla montana (Banks, 1898) Chloroperla - IN, MN, OH, ON
      Isoperla nana (Walsh, 1862) Chloroperla
      Isoperla orata Frison, 1942 - MN, OH
      Isoperla richardsoni Frison, 1935 - Yanoviak and McCafferty 1996:204; Grubbs et al. 2012:166
      Isoperla signata (Banks, 1902) Perlinella (orig.)
        (comb.) Walshiola signata Banks, 1948
        (syn.) Pictetia bimaculata Banks, 1948
      Isoperla slossonae (Banks, 1911) Perla
        (syn.) Clioperla annecta Needham and Claassen, 1925
      Isoperla transmarina (Newman, 1838) Chloroperla (orig.) - Frison 1942, Yanoviak and McCafferty 1996:204
        (syn.) Isoperla ventralis Banks, 1908
        (syn.) Isoperla fumosa Neave, 1933

PTERONARCYIDAE Newman, 1853 - Salmonflies

    Pteronarcys Newman, 1838
      Pteronarcys dorsata (Say, 1823) Sialis (orig.) - Bright coll.; Yanoviak and McCafferty 1996: 204; widespread in state; see also Nelson 2000:32-33
        (syn.) Pteronarcys flavicornis Provancher, 1876
        (syn.) Pteronarcys frigida Gerstaecker, 1873
        (syn.) Pteronarcys insignis (Pictet, 1841)
        (syn.) Pteronarcys labradoriensis Šámal, 1933
        (syn.) Pteronarcys nobilis Hagen, 1861 (syn.); (Frison 1935 INHSB 20:336 has it as P. pictetii)
        (syn.) Pteronarcys rectus Provancher, 1876
        (syn.) Pteronarcys regalis Newman, 1838
        (syn.) Pteronarcys shelfordi Frison, 1934
      Pteronarcys pictetii Hagen, 1873
        (misid.) Pteronarcys nobilis: Smith 1917, Needham and Claassen 1925, Frison 1935

TAENIOPTERYGIDAE Klapálek, 1905 - Taeniopterygid winter stoneflies

    Oemopteryx Klapálek, 1902 (subfamily Brachypterainae Zwick, 1973)
      Oemopteryx glacialis (Newport, 1849) Nemoura (orig.)
        (syn.) Perla chicoutimiensis Provancher, 1878
        (syn.) Taeniopteryx (Oemopteryx) alex Hanson, 1938

    Strophopteryx Frison, 1929 (subfamily Brachypterainae Zwick, 1973)
      Strophopteryx fasciata (Burmeister, 1839) Semlis
        (syn.) Taeniopteryx frigida Hagen, 1861
        (comb.) Tachiopteryx (Strophopteryx) fasciata Frison, 1942
        (comb.) Brachyptera fasciata Harden & Mickel, 1952

    Taeniopteryx Pictet, 1841 (subfamily Taeniopteryginae Klapálek, 1905)
      Taeniopteryx burksi Ricker & Ross, 1968 - Yanoviak and McCafferty 1996: 204
      Taeniopteryx nivalis Fitch, 1847
      Taeniopteryx parvula Banks, 1918

References

    Alexander KD, Stewart KW. 1999. Revision of the genus Suwallia Ricker (Plecoptera: Chloroperlidae). Transactions of the American Entomological Society 125(3):185-250.
    Banks N. 1908. Neuropteroid insects, notes and descriptions. Transactions of the American Entomological Society 34:255-267.
    Baumann RW. 1974. What is Alloperla imbecilla (Say)? Designation of a neotype, and a new Alloperla from eastern North America (Plecoptera: Chloroperlidae). Proceedings of the Biological Society of Washington 87(24):257-264.
    Boumans L, Baumann RW. 2012. Amphinemura palmeni is a valid Holarctic stonefly species (Plecoptera: Nemouridae). Zootaxa 3537: 59-75.
    Claassen PW. 1924. New species of North American Capniidae (Plecoptera). Canadian Entomologist 56: 43-48, 54-57.
    Claassen PW. 1940. A catalogue of the Plecoptera of the world. Memoirs of the Cornell University Agricultural Experimental Station 232:1-235.
    Frison TH. 1935.. The stoneflies, or Plecoptera, of Illinois. Bulletin of the Illinois Natural History Survey 20:281-471.
    Frison TH. 1937. Descriptions of Plecoptera with special reference to the Illinois species. Bulletin of the Illinois Natural History Survey 21:78-99.
    Frison TH. 1942. Studies of North American Plecoptera, with special reference to the fauna of Illinois. Bulletin of the Illinois Natural History Survey 22:235-355.
    Grubbs SA, Bright E. 2001. Arcynopteryx compacta (Plecoptera: Perlodidae), a Holarctic stonefly confirmed from Lake Superior, with a review and first checklist of the stoneflies of Michigan. The Great Lakes Entomologist 34(2):77-84.
    Grubbs SA, Pessino M, DeWalt RE. 2012. Michigan Plecoptera (Stoneflies): distribution patterns and an updated species list. Illiesia 8(18):162-173.
    Grubbs SA, DeWalt RE. 2011. Perlesta ouabache, a new species of stonefly (Plecoptera: Perlidae) from Indiana, USA. Aquatic Insects 33(1): 75-79.
    Grubbs SA, DeWalt RE. 2012. Perlesta ephelida, a new Nearctic stonefly species (Plecoptera, Perlidae). Zookeys 194: 1-15.
    Hanson JF. 1943. Records and descriptions of North American Plecoptera. Part III. Notes on Isogenoides. American Midland Naturalist 29: 657–669.
    Hitchcock SW. 1974. Guide to the Insects of Connecticut. Part VII. The Plecoptera or stoneflies of Connecticut. State Geological and Natural History Survey of Connecticut Bulletin 107:1-262.
    Needham JG, Claassen PW. 1925. A Monograph of the Plecoptera or Stoneflies of America North of Mexico. Entomological Society of America, The Thomas Say Foundation, Lafayette, Indiana: 1-397.
    Nelson CH. 2000. Pteronarcyidae (the Salmonflies), pp. 29-39 in Stark BP, Armitage BJ (eds.), Stoneflies (Plecoptera) of Eastern North America, Volume 1: Pteronarcyidae, Peltoperlidae, and Taeniopterygidae. Ohio Biological Survey Bulletin New Series Volume 14, Number 1. vii + 100p.
    Sandberg JB, Stewart KW. 2003. Continued studies of drumming in North American Plecoptera; Evolutionary implications, pp. 73-81 in E. Gaino (ed.). Research update on Ephemeroptera and Plecoptera. University of Perugia, Perugia, Italy.
    Sandberg JB, Stewart KW. 2005. Holomorphology and Systematics of the Stonefly Genus Isogenoides (Plecoptera: Perlodidae). Transactions of the American Entomological Society 131(3+4): 269-345.
    Stewart KW. 2000. Taeniopterygidae (the Willowflies), pp. 55-88 in Stark BP, Armitage BJ (eds.), Stoneflies (Plecoptera) of Eastern North America, Volume 1: Pteronarcyidae, Peltoperlidae, and Taeniopterygidae. Ohio Biological Survey Bulletin New Series Volume 14, Number 1. vii + 100p.
    Stewart KW, Stark BP. 2002. Nymphs of North American stonefly genera. Second Edition. The Caddis Press: Columbus, Ohio. xii + 510
    Szczytko SW, Kondratieff BC. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia 1:1-289.
    Surdick RF. 1985. Nearctic genera of Chloroperlidae (Plecoptera: Chloroperlidae). Illinois Biological Monographs 54:1-146.
    Teslenko VA. 2012. A taxonomic revision of the genus Arcynopteryx Klapálek, 1904 (Plecoptera, Perlodidae). Zootaxa 3329:1-18.
    USGS Northern Prairie Wildlife Rearch Center - web site managed by B. Kondratieff (accessed August 6, 2006)
    Yanoviak SP, McCafferty WP. 1996. Comparison of macroinvertebrate assemblages inhabiting pristine streams in the Huron Mountains of Michigan, USA. Hydrobiologia 330: 195-211.
Page edits:
    March 23, 2015
    • Isoperla decepta added to the state species list, as listed in a new publication by Szczytko and Kondratieff (2015)
    • Leuctra tenuis, previously listed, removed in Grubbs and Bright (2001) as the record from Yanoviak and McCafferty 1996 (p. 204), based on nymphs, proved unsupported, and its currently known distribution (and in the updated Michigan list by Grubbs et al. (2012) make it unlikely to be recorded in Michigan. Therefore, I have removed this species from the list. I have also removed a number of other species (Allocapnia frisoni, Allocapnia rickeri, Taeniopteryx metequi, Perlesta adena), although regionally recorded, have distributions from only unglaciated areas and are not likely to be found in Michigan.
    March 20, 2015
    • Paranemoura perfecta and Arcynopteryx dichroa, listed in Grubbs et al. (2012), has also been added to the list. This edit supercedes the edit dated November 04, 2012 (see below) for Arcynopteryx compacta, which is now in synonomy with Arcynopteryx dichroa.
    • Leuctra tenuis, previously listed, removed in Grubbs and Bright (2001) as the record from Yanoviak and McCafferty 1996 (p. 204), based on nymphs, proved unsupported, and its currently known distribution (and in the updated Michigan list by Grubbs et al. (2012) make it unlikely to be recorded in Michigan. Therefore, I have removed this species from the list.
    January 04, 2013
    • Taeniopteryx maura, recorded as new to Michigan by Grubbs and Bright (2001), was based on a prior misidentification of T. burksi, and is removed from the list.
    • Isogenoides varians, previously recorded in Michigan by Needham and Claassen (1925: 84) and Hanson (1943), are unsubstantiated without location of specimens or misidentifications; based on it's known distribution, it is unlikely to be found in Michigan (Sandberg and Stewart 2005). It is removed from the list.
    December 21, 2012
    • Boumans & Baumann (2012) recently synonomyzed Amphinemura linda with Amphinemura palmeni based on morphological and genetic evidence
    • Grubbs & DeWalt (2012) described Perlesta ephelida, found in Michigan; previous records of Perlesta shubuta from Michigan (Grubbs and Bright 2001) are of P. ephelida
    November 04, 2012
    • Beginning to keep track of changes to the web site pages
    • Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page
    • Updated synonomies per Plecoptera Home Page
    • Added Perlesta ouabache (Grubbs & DeWalt, 2011).
    • Arcynopteryx compacta (McLachlan, 1872) moved to Skwala Ricker, 1943 (Teslenko 2012).

Page created: October 29, 2002 - Last updated: March 24, 2015 (EB)