Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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CERATOPOGONIDAE Newman, 1834 (No-See-Ums, Biting midges)

Below is a preliminary list of species based on records in the UMMZ-Insect Division collection (identified by Wirth, Stone and Steyskal) as well as literature sources, principally those of Wirth, Borkent and Grogan.

    Allohelea Kieffer, 1917 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Allohelea johannseni (Wirth, 1953)
        (orig.) Monohelea johannseni Wirth, 1953:153

    Alluaudomyia Kieffer, 1913 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Alluaudomyia bella (Coquillett, 1902) - widespread, WI (Borkent and Grogan 2009:18)
        (orig..) Ceratopogon bella Coquillett, 1902: 87
        (syn.) Isoecacta poeyi Garrett, 1925:9
        (syn.) Alluaudomyia splendida, authors, not Winnertz
      Alluaudomyia footei Wirth, 1952 - Borkent and Grogan 2009:18
      Alluaudomyia megaparamera Williams, 1957 - Williams 1957:327; Borkent and Grogan 2009:18
      Alluaudomyia needhami Thomsen, 1935 - widespread, WI (Borkent and Grogan 2009:18)
      Alluaudomyia parva Wirth, 1952 - Williams 1956; see also Grogan and Bystrak 1976
        (syn.) Alluadomyia downesi Wirth, 1952:431
      Alluaudomyia wirthi Williams, 1957 - Williams 1957:328; Borkent and Grogan 2009:18

    Atrichopogon Kieffer, 1906 (Subfamily Forcipomyiinae)

      Atrichopogon (Meloehelea) downesi Wirth, 1980:129 - MN, east (Borkent and Grogan 2009:6)
      Atrichopogon (Meloehelea) epicautae Wirth, 1956:21 - widespread (Borkent and Grogan 2009:6)
      Atrichopogon (Atrichopogon) falcatus Boesel, 1973 - Boesel 1973:214-215 (holotype from Emmet Co.)
      Atrichopogon (Lophomyidium) fusculus Coquillett, 1901) - widespread (Boesel 1973:209; Borkent and Grogan 2009:6; Malloch 1915:304-306)
        (orig.) Ceratopogon fusculus Coquillett, 1901:605
      Atrichopogon (Atrichopogon) fusinervis (Malloch, 1915) - Boesel 1973:211; see also Malloch 1915:308
        (orig.) Ceratopogon fusinervis Malloch, 1915:308
      Atrichopogon (Atrichopogon) geminus Boesel, 1973 - OH, ON (Boesel 1973:211-213)
      Atrichopogon (Atrichopogon) levis (Coquillett, 1901) - UMMZ; Malloch 1915:307-308
        (orig.) Ceratopogon levis Coquillett, 1901:604
        (syn.) Ceratopogon exilis Coquillett, 1902:86
      Atrichopogon (Meloehelea) lucorum (Meigen, 1818) - widespread (Borkent and Grogan 2009:6)
        (orig.) Ceratopogon lucorum Meigen, 1818:72
      Atrichopogon (Atrichopogon) minutus (Meigen, 1830) - OH (Boesel 1973:206-207), widespread (Borkent and Grogan 2009:5)
        (orig.) Ceratopogon minutus Meigen, 1830:263
      Atrichopogon (Atrichopogon) peregrinus (Johannsen, 1908) - UMMZ; Malloch 1915:308-309; see also Boesel 1973:210-211
        (orig.) Ceratopogon peregrinus Johannsen, 1908:266
      Atrichopogon (Atrichopogon) titanus Boesel, 1973 - OH (Boesel 1973:213-214), expect in se LP
      Atrichopogon (Atrichopogon) websteri (Coquillett, 1901) - IL, OH, ON, widespread (Boesel 1973:207-208; Borkent and Grogan 2009:5)
        (orig.) Ceratopogon websteri Coquillett, 1901:603

    Bezzia Kieffer, 1899 (Subfamily Ceratopogoninae, Tribe Palpomyiini)

      Bezzia (Bezzia) albidorsata Malloch, 1915:349 - IL, eastern (Borkent and Grogan 2009:26)
      Bezzia (Bezzia) apicata Malloch, 1914:284 - Borkent and Grogan 2009:18
      Bezzia (Homobezzia) bicolor
        (orig.) Ceratopogon bicolor Meigen, 1804:31
        (syn.) Probezzia copiosa Thomsen, 1935:292
      Bezzia (Bezzia) bivittata (Coquillett, 1905) - Wirth and Grogan 1983: 494,499-501, 503
        (orig.) Ceratopogon bivittatus Coquillett, 1905:60
        (comb.) Probezzia bivittata (Coquillett, 1905), Johannsen 1943:267, Malloch 1914:138
        (comb.) Bezzia (Aspinabezzia) bivittata (Coquillett, 1905), Dow and Turner 1976:126
      Bezzia (Homobezzia) cockerelli Malloch, 1915:346 - WI, widespread (Borkent and Grogran 2009:27)
        (syn.) Probezzia obscura Malloch, 1915:355
      Bezzia (Bezzia) dentata Malloch 1914:284 - IL, widespread (Borkent and Grogan 2009:26)
      Bezzia (Bezzia) expolita (Coquillett, 1901) - widespread (Borkent and Grogan 2009:26)
        (orig.) Ceratopogon exploitus Coquillett, 1901:600
        (syn.) Ceratopogon johnsoni Coquillett, 1901:600
      Bezzia (Bezzia) flavitarsis Malloch, 1914:283 - IL, widespread (Borkent and Grogan 2009:26)
      Bezzia (Bezzia) gibberella Wirth & Grogan, 1983:508 - Wirth and Grogan 1983:508-510
      Bezzia (Homobezzia) glabra (Coquillett, 1902) - UMMZ
        (orig.) Ceratopogon glabrus Coquillett, 1902:85
      Bezzia (Bezzia) nobilis (Winnertz, 1852) - Widespread (Borkent and Grogan 2009:26)
        (orig.) Ceratopogon nobilis Winnertz, 1852:79
        (syn.) Bezzia atlantica Wirth and Williams, 1957:13
        (syn.) Ceratopogon barberi Coquillett, 1901:601
        (syn.) Ceratopogon setulosa Loew, 1861:312
      Bezzia (Homobezzia) obelisca Dow & Turner, 1976:104 - Borkent and Grogan 2009:27
      Bezzia (Bezzia) uncistyla Dow & Turner, 1976:80 - widespread (Borkent and Grogan 2009:26)
      Bezzia (Homobezzia) varicolor (Coquillett, 1902) - ON, WI, widespread (Borkent and Grogan 2009:27)

    Brachypogon Kieffer, 1899 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Brachypogon canadensis Downes, 1976:1147 - Cooper 1991:9; see also Borkent and Grogan 1995:46
      Ceratopogon multisetosus Borkent and Grogan, 1995:76 - NB, BC, widespread northern (Borkent and Grogan 1995:76)
      Ceratopogon pandus Borkent and Grogan, 1995:53 - WI (Borkent and Grogan 1995:46)

    Ceratopogon Meigen, 1803 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Ceratopogon inverecundus Borkent and Grogan, 1995:64 - widespread northern (Borkent and Grogan 1995:46)
      Ceratopogon multisetosus Borkent and Grogan, 1995:76 - NB, BC, widespread northern (Borkent and Grogan 1995:76)
      Ceratopogon pandus Borkent and Grogan, 1995:53 - WI (Borkent and Grogan 1995:46)

    Clinohelea Kieffer, 1917 (Subfamily Ceratopogoninae, Tribe Heteromyiini)

      Clinohelea bimaculata (Loew, 1861) - Grogan and Wirth 1975:281-283
        (orig.) Ceratopgon bimaculatus Loew, 1861:311
        (comb.) Johannseniella bimaculata (Loew, 1861), Malloch 1914:226
        (comb.) Johannsenomyia bimaculata (Loew, 1861), Malloch 1915:332
      Clinohelea curriei (Coquillett, 1905) - Grogan and Wirth 1975:277-279; Malloch 1915:322 (as P. nebulosa)
        (orig.) Ceratopogon curriei Coquillett, 1905:62
        (comb.) Palpomyia curriei (Coquillett, 1905); Malloch 1914:219
        (comb.) Clinohelea curriei (Coquillet, 1905); Johannsen 1943: 783
        (syn.) Palpomyia nebulosa Malloch, 1915:322, Grogan and Wirth 1975:278
        (syn.) Clinohelea nebulosa (Malloch, 1915), Johannsen 1943:783

    Culicoides Latreille, 1809 (Subfamily Ceratopogoninae, Tribe Culicoidini)

      Culicoides (unplaced) alexanderi Wirth and Hubert, 1962 - Borkent and Grogan 2009:16
        (syn.) Culicoides pseudoptiliferus Wirth & Hubert, 1962:189
      Culicoides (Amossovia) arboricola Root & Hoffman, 1937:166 - ON, MN, WI, widespread (Borkent and Grogan 2009:12)
      Culicoides (Diphaomyia) baueri Hoffman, 1925 - WI west and east
      Culicoides (Diphaomyia) bergi Cochrane, 1973 - WI, WY-NY, south (Cochrane, 1973:311; Borkent and Grogan 2009:13)
      Culicoides (unplaced) bickleyi Wirth & Hubert, 1962 - WI east and south
      Culicoides (Oecacta) biguttatus (Coquillett, 1901) Ceratopogon (orig.) - UMMZ
      Culicoides (Avaritia) chiopterus (Meigen, 1830) - widespread (Borkent and Grogan 2009:12)
        (orig.) Ceratopogon chiopterus Meigen, 1830:263
      Culicoides (Beltranmyia) crepuscularis Malloch, 1915:303-304 - IL west and east (Borkent and Grogan 2009:13)
      Culicoides (unplaced) denticulatuts Wirth & Hubert, 1962:193 - MN, ON, WI, widespread (Borkent and Grogan 2009:16)
        (syn.) Culicoides unicorlor, authors, not Coquillett
      Culicoides (unplaced) downesi Wirth & Hubert, 1962:186 - Borkent and Grogan 2009:17
      Culicoides (Amossovia) flukei Jones, 1956:30 - ON, WI, east (Borkent and Grogan 2009:12)
      Culicoides (Oecacta) furensoides Williams, 1955:271 - Borkent and Grogan 2009:15
        (syn.) Culicoides dickei Jones, 1956:28
      Culicoides (Amossovia) guttipennis (Coquillett, 1901) - MN, OH (Borkent and Grogan 2009:12)
        (orig.) Ceratopogon guttipennis Coquillett 1901:603
      Culicoides (Diphaomyia) haematopotus Malloch, 1915:302-303 - IL west and east (Borkent and Grogan 2009:13)
      Culicoides (unplaced) jamnbacki Wirth & Hubert, 1962:192 - Borkent and Grogan 2009:17
      Culicoides (unplaced) nanus Root & Hoffman, 1937:165 - ON, WI; see also Borkent and Grogan 2009:17
      Culicoides (Avaritia) obsoletus (Meigen, 1818) - widespread (Borkent and Grogan 2009:12)
        (orig.) Ceratopogon obsoletus Meigen, 18198:76
      Culicoides (Haematomyidium) paraensis (Goeldi, 1905) - WI, CO-PA (Borkent and Grogan 2009:14)
        (orig.) Haematomyidium paraensis Goeldi, 1905
      Culicoides (Culicoides) paraimpunctatus Borkent, 1995 - MN, WI, widespread north (Borkent and Grogan 2009:13)
        (syn.) Culicoides canadensis Wirth and Blanton, 1969:211 (preocc. by Boese, 1937)
      Culicoides (unplaced) piliferus Root & Hoffman, 1937 - UMMZ; see also Borkent and Grogan 2009:17
      Culicoides (Avaritia) sanguisuga (Coquillett, 1901) - widespread (Borkent and Grogan 2009:12)
        (orig.) Ceratopogon sanguisuga Coquillett, 1901:604
        (syn.) Simulium nocivum Harris, 1862
      Culicoides (unplaced) scanloni Wirth & Hubert, 1962:187 - Borkent and Grogan 2009:17
      Culicoides (Beltranmyia) sphagnumensis Williams, 1955:269 - see also Cooper 1991:11 (Paratypes)
      Culicoides (Silvaticulicoides) spinosus Root & Hoffman, 1937:172 - ON, WI (Borkent and Grogan 2009:15)
      Culicoides (Oecacta) stellifer (Coquillett, 1901) - Borkent and Grogan 2009:15
        (orig.) Ceratopogon stellifer Coquillett, 1901:603
      Culicoides (Wirthomyia) stilobezzioides Foote & Pratt, 1954:33 - AK-NF, MN, south (Borkent and Grogan 2009:16)
      Culicoides (unplaced) travisi Vargas, 1949 - ON, WI, widespread (Borkent and Grogan 2009:17)
        (syn.) Culicoides simulans Root & Hoffman, 1937
        (syn.) Culicoides horneae Foote & Pratt, 1954
      Culicoides (Monoculicoides) variipennis (Coquillett, 1901) - Wirth and Jones 1957
      [Note: this species has been implicated in transmitting epizootic hemorrhagic disease in white-tailed deer (Odocoileus virginianus)
        (orig.) Ceratopogon varipennis Coquillett, 1902:94
        (spell.) Culicoides varipennis (Coquillett, 1902), Malloch 1915:297
      Culicoides (Hoffmania) venustus Hoffman, 1925 - widespread east (Borkent and Grogan 2009:14)
      Culicoides (Amossovia) villosipennis Root & Hoffman, 1937:165 - WI southwest and east (Borkent and Grogan 2009:12)
      Culicoides (Beltranmyia) wisconsinensis Jones, 1956:32 - see also Borkent and Grogan 2009:13

    Dasyhelea Kieffer, 1911 (Subfamily Dasyheleinae)

      Dasyhelea ampla Borkent, in Borkent & Wirth, 1997:51 - Waugh and Wirth 1976:224-225 (as D. major)
        (syn.) Pscudoculicoides major Malloch, 1915:311; preocc. by Stroble, 1906
        (syn.) Dasyhelea major (Malloch, 1915), Thomsen, 1935:23
      Dasyhelea cincta (Coquillett, 1901) - Malloch 1915:311 (42.405°N,-86.042°W [estm.])
        (orig.) Ceratopogon cinctus Coquillett, 1901:605
        (comb.) Culicoides cinctus (Coquillett, 1901), Kieffer 1906:54
        (comb.) Pseudoculicoides cinctus (Coquillett, 1901), Malloch 1915:311
        (comb.) Dasyhelea cincta (Coquillett, 1901), Johannsen, 1943:788; Wirth 1952:150
      Dasyhelea flavifrons (Guérin, 1833) - Waugh and Wirth 1976:230-231 (as D. opressa)
        (orig.) Ceratopogon flavifrons Guérin, 1833:165
        (syn.) Dasyhelea opressa Thomsen, 1935:285; Thomsen 1937:68
      Dasyhelea grisea (Coquillett, 1901) - Waugh and Wirth 1976:228-229
        (orig.) Ceratopogon griseus Coquillett, 1901:602
        (comb.) Culicoides griseus (Coquillett, 1901), Kieffer 1906:54
        (comb.) Dasyhelea grisea (Coquillett, 1901), Thomsen 1935:283, Thomsen 1937:37; Wirth 1952:155
        (syn.) Dasyhelea subcaerulea Thomsen, 1935:284
      Dasyhelea leptobranchia Waugh and Wirth, 1976:236 - WI-NY (Borkent and Grogan 2009:11)
      Dasyhelea mutabilis (Coquillett, 1901) - widespread (Malloch 1915:310; Borkent and Grogan 2009:11)
        (orig.) Ceratopogon mutabilis Coquillett, 1901:604
        (comb.) Pseudoculicoides mutabilis (Coquillett, 1901), Malloch 1915:310
      Dasyhelea navaiae Waugh & Wirth, 1976 - Waugh and Wirth 1976:240 Dasyhelea stemlerae Waugh & Wirth, 1976:233 - Borkent and Grogan 2009:11
      Dasyhelea traverae Thomsen, 1935
        (orig.) Dasyhelea traverae Thomsen, 1935:285; Thomsen 1937:68

    Echinohelea Macfie, 1940 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Echinohelea lanei Wirth 1951:319 - Borkent and Grogan 2009:20

    Forcipomyia Meigen, 1818 (Subfamily Forcipomyiinae)

      Forcipomyia (Synthryridomyia) acidicola (Tokunaga, 1937) - widespread (Borkent and Grogan 2009:9)
        (orig.) Lasiohelea acidicola Tokunaga, 1937:455
        (syn.) Forcipomyia colemani Wirth, 1952:146
      Forcipomyia (Forcipomyia) aurea Malloch, 1915:318 - IL
      Forcipomyia (Forcipomyia) bipunctata (Linnaeus, 1767) - widespread, Holarctic (Borkent and Grogan 2009:7)
        (orig.) Tipula bipunctata Linnaeus, 1767:978
      Forcipomyia (Forcipomyia) brevipennis (Macquart, 1826) - BC-QB, s. to LA (Borkent and Grogan 2009:7)
        (orig.) Ceratopogon brevipennis Macquart, 1826:179
        (syn.) Ceratopogon specularis Coquillett, 1901:601
        (syn.) Forcipomyia specularis (Coquillett, 1901)
      Forcipomyia (Forcipomyia) bystraki Grogan & Wirth, 1975 - Grogan and Wirth 1975a:466
      Forcipomyia (Forcipomyia) cilipes (Coquillett, 1900) - widespread (Borkent and Grogan 2009:7)
        (orig.) Ceratopogon cilipes Coquillett, 1900:397
      Forcipomyia (Forcipomyia) concolor Malloch, 1915 - IL (Malloch 1915:319)
      Forcipomyia (Trichohelea) crinita Saunders, 1964:474 - widespread (Borkent and Grogan 2009:10)
      Forcipomyia (Forcipomyia) elegantula Malloch, 1915 - IL, widespread (Grogan and Wirth 1975b: 95-101; see also Wirth and Grogan 1978)
      Forcipomyia (Trichohelea) eques (Johannsen, 1908)
        (orig.) Ceratopogon eques Johannsen, 1908:266
      Forcipomyia (Lasiohelea) fairfaxensis Wirth, 1951:317 - Borketn and Grogan 2009:8
      Forcipomyia (Thyridomyia) frutetorum (Winnertz, 1852) - Borketn and Grogan 2009:9-10
        (orig.) Ceratopogon frutetorum Winnertz, 1852:29
        (syn.) Forcipomyia aspinosa Saunders, 1957:692
      Forcipomyia (Microhelea) fuliginosa (Meigen, 1818) Ceratopogon (orig.) - cosmopolitan, Nearctic widespread (Borkent and Grogan 2009:8)
        (syn.) Forcipomyia erucicida Knab, 1914:65
        (syn.) Ceratopogon coquilletti Kieffer, 1917:297
        (syn.) Focipomyia brookmani Wirth, 1952:140
      Forcipomyia (Caloforcipomyia) glauca Macfie, 1934:144 - Wirth, 1951:315
        (syn.) Forcipomyia splendida Wirth, 1951:315
      Forcipomyia (Forcipomyia) hirtula (Zetterstedt, 1838) - AK-ON, south (Borkent and Grogan 2009:7)
        (orig.) Chironomus hirtulus Zetterstedt, 1838:815
        (syn.) Forcipomyia simulata Walley, 1932:165
      Forcipomyia (Caloforcipomyia) glauca Macfie, 1934:144 - widespread (Borkent and Grogan 2009:6)
      Forcipomyia (Trichohelea) mcateei Wirth, 1956:359
        (syn.) Forcipomyia saundersi Chan, in Chan and LeRoux, 1965:87
      Forcipomyia (Thyridomyia) monilicornis (Coquillett, 1905) - widespread
        (orig.) Ceratopogon monilicornis Coquillett, 1905:63
        (syn.) Thyridomyia palustris Saunders, 1925:269
      Forcipomyia (Euprojoannisia) palustris (Meigen, 1804) - IL, widespread (Borkent and Grogan 2009:7)
        (orig.) Ceratopogon palustris Meigen, 1804:28
        (syn.) Ceratopogon hirtipennis Malloch, 1915:313
        (syn.) Forcipomyia (Euforcipomyia) hirtipennis (Malloch, 1915)
      Forcipomyia (Metaforcipomyia) pluvialis Malloch, 1923:5 - ON south (Borkent and Grogan 2009:8)
      Forcipomyia (Forcipomyia) squamipes (Coquillett, 1902) - widespread (Borkent and Grogan 2009:7)
        (orig.) Ceratopogon squamipes Coquillett, 1902:88
        (syn.) Ceratopogon brumalis Long, 1902:10
      Forcipomyia (Focipomyia) texana (Long, 1902) - widespread
        (orig.) Ceratopogon texana Long, 1902:10

    Heteromyia Say, 1825 (Subfamily Ceratopogoninae, Tribe Heteromyiini)

      Heteromyia prattii Coquillett, 1902:88 - Wirth and Grogan 1977:179-181

    Jenkinshelea Macfie, 1934 (Subfamily Ceratopogoninae, Tribe Spaheromiini)

      Jenkinshelea albaria (Coquillett, 1895) - IL, ON, south and east (Grogan and Wirth 1977:129-135)
        (orig.) Ceratopogon albaria Coquillett, 1895:308
        (syn.) Johannsenomyia aequalis Malloch, 1915:336
      Jenkinshelea magnipennis (Johannsen, 1908) - Grogan and Wirth 1977:134,137-139
        (orig.) Johannseniella magnipennis Johannsen, 1908:268

    Johannsenomyia Malloch, 1915 (Subfamily Ceratopogoninae, Tribe Sphaeromiini)

      Johannsenomyia annulicornis Malloch, 1918 - Wirth 1962:276; Borkent and Grogan 2009:24
        (comb.) Dicrohelea annulicornis (Malloch, 1918), Johannsen 1943:783
      Johannsenomyia argentata (Loew, 1861) - Wirth 1962:277
        (orig.) Ceratopogon argentata Loew, 1861:310
        (comb.) Johannseniella argentata (Loew, 1861), Malloch 1914:226, Malloch 1915:334
        (comb.) Spaeromias argentata (Loew, 1861), Kieffer 1917:315
        (syn.) Ceratopogon halteralis Malloch, 1915:338 (in part)

    Leptoconops Skuse, 1889 (Subfamily Leptoconopinae)

      Leptoconops (Holoconops) catawbae (Boesel, 1948)
        (orig.) Holoconops catawbae Boesel, 1948:69

    Mallochohelea Wirth, 1962 (Subfamily Ceratopogoninae, Tribe Sphaeromiini)

      Mallochohelea albibasis (Malloch, 1915) - Wirth 1962:279-280
        (orig.) Johannsenomyia albibasis Malloch, 1915:315
      Mallochohelea albihalter Wirth, 1962 - Wirth 1962:280
      Mallochohelea atripes Wirth, 1962 - Wirth 1962:281
      Mallochohelea flavidula (Malloch, 1914) - IL, WI (Wirth 1962:281-282; Borkent and Grogan 2009:24)
        (orig.) Johannsenomyia flavidula Malloch, 1914:230
      Mallochohelea smithi (Lewis, 1956) - Wirth 1962:282
        (orig.) Johannsenomyia smithi Lewis, 1956:47
      Mallochohelea variegata Wirth, 1962:283 - Wirth 1962:283

    Monohelea Kieffer, 1917 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Monohelea grogani Delécolle & Rieb, 1995 - Borkent and Grogan 2009:20
      Monohelea jamnbacki Wirth & Williams, 1964 - Wirth & Williams, 1964:308
      Monohelea macflei Wirth, 1953:143
      Monohelea magnitheca Wirth & Grogan, 1981:42 - ON, WI, widespread north (Borkent and Grogan 2009:20)
      Monohelea obscura Wirth & Williams, 1964:304 - Borkent and Grogan 2009:20

    Nilobezzia Kieffer, 1921 (Subfamily Ceratopogoninae, Tribe Spaheromiini)

      Nilobezzia mallochi Wirth, 1962 - Wirth 1962:284-285
      Nilobezzia minor (Wirth, 1952) - Wirth 1962:285; see also Borkent and Grogan 2009:24
        (orig.) Spaheromias minor Wirth, 1952:216

    Palpomyia Meigen, 1818 (Subfamily Ceratopogoninae, Tribe Palpomyiini)

      Palpomyia altispina Grogan & Wirth, 1975 - WI, east (Grogan and Wirth 1979:102-106; see also Grogan and Wirth 1975:32-33
      Palpomyia basalis (Walker, 1848) - Grogan and Wirth 1975:27-29; Grogan and Wirth 1979:90-95
        (orig.) Ceratopogon basalis Walker, 1848:27
        (syn.) Ceratopogon trivialis Loew, 1861:309
        (syn.) Palpomyia trivialis (Loew, 1861), Malloch 1914:217
        (syn.) Heteromyia trivialis (Loew, 1861), Malloch 1915:320
        (syn.) Heteromyia tenuicornis Malloch, 1915:328
        (syn.) Palpomyia tenuicornis (Malloch, 1915)
        (misid.) Palpmyia flavipes (Meigen, 1804), Johannsen 1943:784
        (misid.) Palpomyia praeusta (Loew, 1869), Wirth 1965:140
      Palpomyia canadensis Grogan & Wirth, 1979 - widespread (Grogan and Wirth 1979:109-112)
      Palpomyia downesi Grogan & Wirth, 1979 - AK-MA, widespread north (Grogan and Wirth 1979:112-115)
      Palpomyia flaviceps (Johannsen, 1905) - Grogan and Wirth 1975:30-31; Grogan and Wirth 1979:106-109
        (orig.) Ceratolophus flaviceps Johannsen, 1905:105
        (comb.) Johannseniella flaviceps Johannsen, 1908
        (syn.) Palpomyia flaviceps (Johannsen, 1908), syn. of P. flavipes (Meigen, 1804) Johannsen 1943:784
        (syn.) Heteromyia opacithroax Malloch, 1915:329
        (syn.) Palpomyia opacithorax (Malloch, 1915), Johannsen 1943:784
        (syn.) Heteromyia hirta Malloch, 1915:330
        (syn.) Palpomyia hirta (Malloch, 1915), Johannsen 1943:784
      Palpomyia hastata Grogan & Wirth, 1975 - Grogan and Wirth 1975:34-36; however, see Grogan and Wirth 1979:95-98
      Palpomyia jonesi Grogan & Wirth, 1975 - Grogan and Wirth, 1975:17-18; Grogan and Wirth 1979:45-48
      Palpomyia lineata (Meigen, 1818) - Grogan and Wirth 1975:15-17; Grogan and Wirth 1979:39-45
        (orig.) Ceratopogon lineatus Meigen, 1818:80
        (syn.) Ceratopogon slossonae Coquillett, 1905:61
        (syn.) Ceratopogon cinerea Macquart, 1826: 177
        (syn.) Ceratopogon binotata Staeger, 1839: 596
        (syn.) Palpomyia slossonae (Coquillett, 1905), of authors
        (syn.) Palpomyia illlinoisensis Malloch, 1914: 219
        (syn.) Palpomyia pruinescens Thomsen, 1935:290
        (syn.) Palpomyia algarum Kieffer, in Kieffer and Thienemann, 1908: 1
        (syn.) Palpomyia winnertzi Kieffer, 1914a: 238
        (syn.) Palpomyia octasema Kieffer, 1914a: 239
        (syn.) Palpomyia spinosissima Kieffer, 1915c: 285
        (syn.) Palpomyia unifasciata Kieffer, 1922e: 72
        (syn.) Palpomyia murina Kieffer, 1925a: 426
        (syn.) Palpomyia obesa Goetghebuer, 1927b: 94
        (syn.) Palpomyia lineata var nigrotibialis Goetghebuer, 1933d: 370
        (syn.) Palpomyia pruinescens Thomsen, 1935: 290
        (syn.) Palpomyia edwardsi de Meijere, 1946: 10 (preoccupied by Palpomyia edwardsi Cockerell, 1921
        (syn.) Palpomyia stagnalis Clastrier, 1962b: 274
      Palpomyia novitibialis Grogan & Wirth, 1975 - Grogan and Wirth 1979:31-35; see also Grogan and Wirth 1975:13-14
      Palpomyia plebeiella Grogan & Wirth, 1975 - Grogan and Wirth 1975:20-22; Grogan and Wirth 1979:60-62
      Palpomyia plebeia (Loew, 1861) - Grogan and Wirth 1979:56-60
        (orig.) Ceratopogon plebius Loew, 1861:313
        (comb.) Heteromyia plebeia (Loew, 1861), Malloch 1915:327
      Palpomyia rubiginosa Grogan & Wirth, 1975 - Grogan and Wirth 1975:22-24; Grogan and Wirth 1979:64-67
      Palpomyia rufa (Loew, 1861) - Grogan and Wirth 1975:25-26; Grogan and Wirth 1979
        (orig.) Ceratopogon rufus Loew, 1861:314
        (comb.) Heteromyia rufa (Loew, 1861), Malloch 1915:325
        (misid.) Palpomyia rubiginosa Grogan & Wirth, 1975:22 (male allotype)
      Palpomyia scalpellifera Grogan & Wirth, 1975 - Grogan and Wirth 1975:33-34; Grogan and Wirth 1979;99-102
      Palpomyia subaspera (Coquillett, 1901) - Grogan and Wirth 1975c:10-12; Grogan and Wirth 1979:23-26
        (orig.) Ceratopogon subasper Coquillett, 1901:606
        (syn.) Palpmyia essigi Wirth, 1952:225
      Palpomyia tibialis (Meigen, 1818) - Grogan and Wirth 1979:29-31
        (orig.) Ceratopogon tibialis Meigen, 1818:82

    Parabezzia Malloch, 1915 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Parabezzia petiolata Malloch, 1915 - IL, may occur in sw LP (Grogan and Wirth 1977:78-79)
      Parabezzia williamsi Wirth 1965:229 - Grogan and Wirth 1977:64-65, holotype from Douglas Lake, Cheboygan Co.

    Phaenobezzia Haeselbarth, 1965 (Subfamily Ceratopogoninae, Tribe Palpomyiini)

      Phaenobezzia fulvithorax (Malloch, 1915) - Wirth and Grogan 1982:185-189; Cooper 1991:18 (Paralectotype)
        (orig.) Probezzia fulvithorax Malloch, 1915:354
        (comb.) Bezzia fulvithorax (Malloch, 1915), Johannsen, 1952:167 (in key)
        (syn.) Bezzia opaca (Loew, 1861), in Wirth 1952:227
      Phaenobezzia opaca (Loew, 1861) - Wirth and Grogan 1982:181-185
        (orig.) Ceratopogon opacus Loew, 1861:321
        (comb.) Bezzia opaca (Loew, 1861), Kieffer, 1906:58
        (comb.) Bezzia (Probezzia) opaca (Loew, 1861), Kieffer 1917:329
        (comb.) Probezzia opaca (Loew, 1861), Malloch 1914:139
        (comb.) Phaenobezzia opaca (Loew, 1861), Haeselbarth 1965:302
        (syn.) Probezzia incerta Malloch, 1915:358; Johannsen 1943:785
        (syn.) Bezzia incerta (Malloch, 1915), Johannsen, 1952:167

    Probezzia Kieffer, 1906 (Subfamily Ceratopogoninae, Tribe Palpomyiini)

      Probezzia albitibia Wirth, 1971:732 - ON, WI south and east (Borkent and Grogan 2009:25)
        (orig.) Ceratopogon albiventris Loew, 1861:311
      Probezzia atriventris Wirth, 1951 - Wirth 1951:31
      Probezzia concinna (Meigen, 1818) - QB, ON west
        (orig.) Ceratopogon concinna Meigen, 1818:77
      Probezzia infuscata Malloch, 1915:316 - IL, MN, ON (Borkent and Grogan 2009:25)
      Probezzia jambacki Wirth, 1971 - Wirth, 1971:735
      Probezzia pallida Malloch, 1914:138 - IL, ON, WI (Borkent and Grogan 2009:25)
      Probezzia sabroskyi Wirth, 1951 - Wirth, 1951:31
      Probezzia smithii (Coquillett, 1901) - WI east and south (Borkent and Grogan 2009:25)
        (orig.) Ceratopogon smithii Coquillett, 1901:600
      Probezzia williamsi Wirth 1971 - Wirth 1971:737
      Probezzia xanthogaster (Kieffer, 1917) - IL, ON, WI (Borkent and Grogan 2009:25)
        (orig.) Bezzia xanthogaster Kieffer, 1917:329
        (syn.) Probezzia elegans Coquillett, 1901:599

    Schizohelea Kieffer, 1917 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Schizohelea leucopeza (Meigen, 1804) - IL, widespread (Borkent and Grogan 2009:21)
        (orig.) Ceratopogon leucopexa Meigen, 1804:29
        (syn.) Ceratopogon politus Colquillett, 1901:606

    Serromyia Meigen, 1818 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Serromyia borealis Borkent, 1990 - Borkent and Bissett 1990:180,186-187
      Serromyia crassifermorata Malloch, 1914 - Borkent and Bissett 1990:180,187
      Serromyia ledicola Kieffer 1925 - ON, AK-VA widespread (Borkent and Bissett 1990:181,192-194)
      Serromyia nudicolis Borkent 1990 - Borkent and Bissett 1990:180,188-190
      Serromyia vockerothi Borkent 1990 - MN, widespread north (Borkent and Bissett 1990:180,191-192)

    Sphaeromias Curtis, 1829 (Subfamily Ceratopogoninae, Tribe Sphaeromiini)

      Sphaeromias longipennis (Loew, 1861) - Wirth 1962:278; see also Borkent and Grogan 2009:25
        (orig.) Ceratopogon longipennis Loew, 1861:313
        (comb.) Palpomyia (Sphaeromyas) longipennis (Loew, 1861), Kieffer 1906:62
        (comb.) Palpomyia longipennis (Loew, 1861), of authors
        (comb.) Homohelea longipennis (Loew, 1861), Kieffer 1917:364
        (comb.) Sphaeromias longipennis (Loew, 1861), Johannsen 1943:784, Wirth 1952:214

    Stilobezzia Kieffer, 1911 (Subfamily Ceratopogoninae, Tribe Ceratopogonini)

      Stilobezzia (Stilobezzia) antennalis (Coquillett, 1901) - widespread north, ON
        (orig.) Ceratopogon antennalis Coquillett, 1901:606
      Stilobezzia (Stilobezzia) bulla Thomsen, 1935:289 - BC-NB, south (Borkent and Grogan 2009:22)
      Stilobezzia (Stilobezzia) coquilletti Kieffer, 1917:308 - IL, may occur in s LP (Borkent and Grogan 2009:22)
        (syn.) Ceratopogon picta (Coquillett, 1905:60), preocc. by Meigen, 1818
      Stilobezzia (Acanthohelea) lutea (Malloch, 1918) - IL, ON east and south (Borkent and Grogan 2009:22)
        (orig.) Hartomyia lutea Malloch, 1918:18
        (syn.) Stilobezzia mallochi Hoffman, 1924
      Stilobezzia (Stilobezzia) sybleae Wirth, 1953:82 - Borkent and Grogan 2009:22

References

    Boesel MW. 1973. The genus Atrichopogon (Diptera, Ceratopogonidae) in Ohio and neighboring states. The Ohio Journal of Science 73(4):202-215.
    Borkent A, Bissett B. 1990. A revision of the Holarctic species of Serromyia Meigen (Diptera: Ceratopogonidae). Systematic Entomology 15(2):153-217.
    Borkent A, Grogan WL. 2009. Catalog of the New World biting midges north of Mexico (Diptera: Ceratopogonidae). Zootaxa 2273:1-48.
    Cochrane AH. 1973 Two new Nearctic species of Culicoides (Diptera: Ceratopogonidae). The Florida Entomologist 56(4):311-318.
    Cooper B. 1991. Diptera types in the Canadian National Collection. Part 1: Nematocera. Research Branch Agriculture Canada Publication 1845/B. 113 p.
    Coquillett DW. 1901. New Diptera in the U. S. National Museum. Proceedings of the U.S. National Museum 23:593-618.
    Grogan WL, Bystrak PG. 1976. Description of the pupa, larva, and larval habitat of Alluaudomyia parva Wirth (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 78(3):314-316.
    Grogan WL, Wirth WW. 1975a. A new Nearctic species of Forcipomyia (Forcipomyia) described in all stages (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 77(4):466-471.
    Grogan WL, Wirth WW. 1975b. Notes on the systematics and biology of the biting midge, Forcipomyia elegantula Malloch (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 80(1):94-102.
    Grogan WL, Wirth WW. 1975c. A revision of the genus Palpomyia Meigen of northeastern North America (Diptera: Ceratopogonidae). Agriculture Experiment Station, University of Maryland, Contribution Number 5076, August, 1975. 49 pp.
    Grogan WL, Wirth WW. 1977. A revision of the Nearctic species of Jenkinshelea MacFie (Diptera:Ceratopogonidae). Proceedings of the Entomological Society of Washington 79(1):126-141.
    Grogan WL, Wirth WW. 1977. A revision of the Nearctic species of Parabezzia Malloch (Diptera:Ceratopogonidae). Journal of the Kansas Entomological Society 50:49-83.
    Grogan WL, Wirth WW. 1979. The North American predaceous midges of the genus Palpomyia Meigen (Diptera:Ceratopogonidae). Memoirs of the Entomological Society of Washington 8: vi + 125 pp.
    Hoffmann WA. 1924. Stilobezzia mallochi and Atrichopogon gilva (Dipt.: Chironomidae). Entomological News 35:282–284.
    Jamnback H, Wirth WW. 1963. The species of Culicoides related to obsoletus in eastern North America (Diptera: Ceratopogonidae). Annals of the Entomological Society of America 56:185–198.
    Johannsen OA. 1907. Some new species of Kansas Chironomidae, pp. 109–112 in Tucker ES. Some results of desultory collecting of insects in Kansas and Colorado. Bulletin of the University of Kansas, Science Bulletin 4:51–112.
    Johannsen OA. 1908. New North American Chironomidae, pp. 264–285 in Felt EP. 23rd report of the State Entomologist on injurious and other insects of the State of New York, 1907. New York State Museum, Museum Bulletin 124:5–541, 44 pls.
    Johannsen OA. 1934. New species of North American Ceratopogonidae and Chironomidae. Journal of the New York Entomological Society 42:343–352.
    Johannsen OA. 1943. Two new species of American Ceratopogonidae (Diptera). Annals of the Entomological Society of America 36:761–762.
    Kieffer JJ. 1917. Chironomides d'Amérique conservés au Musée National Hongrois de Budapest. Annales Historico-Naturales Musei Nationalis Hungarici 15:292–364.
    Malloch JR. 1915. The Chironomidae, or midges, of Illinois, with particular reference to the species occurring in the Illinois River. Bulletin of the Illinois State Laboratory of Natural History 10(6):273-543, 40 pl.
    Meigen JW. 1804. Klassifikazion und Beschreibung der europäischen zweiflügeligen Insekten. (Diptera Linn.). Erster Band. Abt. I. pp. i–xxviii + 1–152, pls. 1–8; Abt. II, pp. i–vi + 153–314, pls. 9–15. Braunschweig.
    Meigen JW. 1818. Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Vol. 1, xxxvi + 333 pp., pls. 1–11. Aachen.
    Thomsen L. 1935. New species of New York State Ceratopogonidae. Journal of the New York Entomological Society 43:283–297.
    Waugh WT, Wirth WW. 1976. A revision of the genus Dasyhelea Kieffer of the Eastern United States north of Florida (Diptera:Ceratopogonidae). Annals of the Entomological Society of America 69(2):219-247.
    Williams RW. 1955. Two new species of Culicoides from Cheboygan County, Michigan (Diptera, Heleidae). Proceedings of the Entomological Society of Washington 57:269–274.
    Williams RW. 1957.
    Two new species of Alluaudomyia from Cheboygan County, Michigan, with a note on the synonymy of parva and downesi (Diptera: Heleidae). Proceedings of the Entomological Society of Washington 58(6):327-331. note: original title has "para" [sic]
    Wirth WW. 1952. The genus Alluaudomyia Kieffer in North America (Diptera, Heleidae). Annals of the Entomological Society of America 45(3):423-434.
    Wirth WW. 1953. Biting midges of the heleid genus Stilobezzia in North America. Proceedings of the United States National Museum 103:57–85.
    Wirth WW. 1962. A reclassification of the Palpomyia-Bezzia-Macropeza groups, and a revision of the North American Sphaeromiini (Diptera, Ceratopogonidae). Annals of the Entomological Society of America 55:272–287.
    Wirth WW. 1965. Family Ceratopogonidae (Heleidae). Pp. 121-142 in A Catalog of the Diptera of America North of Mexico. Stone A, Sabrosky CW, Wirth WW, Foote RH, Coulson JR. (Editors). Agricultural Research Service, Agricultural Handbook 267, United States Department of Agriculture. Smithsonian Institution Press 1983 reprint: Washington, D. C. iv + 1696.
    Wirth WW. 1971. Six new North American species of Probezzia (Diptera: Ceratopogonidae), with biological notes and a key to species. Annals of the Entomological Society of America 64:729–739.
    Wirth WW, Grogan WL. 1977. Taxonomic notes on the genus Heteromyia Say, and a new species from Nicaragua (Diptera: Ceratopogonidae). The Florida Entomologist 60(3):177-185.
    Wirth WW, Grogan WL. 1978. Notes on the systematics and biology of the biting midge, Forcipomyia elegantula Malloch (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 80(1):94-102.
    Wirth WW, Grogan WL. 1982. The predaceous midges of the genus Phaenobezzia in North America (Diptera: Ceratopogonidae). Memoirs of the Entomological Society of Washington 1982(10):179-192.
    Wirth WW, Grogan WL. 1983. The Nearctic species of the Bezzia bivittata group (Diptera: Ceratopogonidae). Proceedings of the Biological Society of Washington 96(3):489-523.
    Wirth WW, Hubert AA. 1962. The species of Culicoides related to piliferus Root and Hoffman in eastern North America (Diptera, Ceratopogonidae). Annals of the Entomological Society of America 55:182–195.
    Wirth WW, Jones RH. 1957. The North American subspecies of Culicoides variipennis (Diptera, Heleidae). United States Department of Agriculture Technical Bulletin 1170: 1-35.
    Wirth WW, Williams RW. 1964. New species and records of North American Monohelea (Diptera: Ceratopogonidae). Annals of the Entomological Society of America 57:302–310.
Page edits:
    February 07, 2013
    • Incorporated nomenclature of Borkent and Grogan 2009, including some new state records
    November 21, 2012
    • Beginning to keep track of changes to the web site pages
    • Ceratopogonidae retained as a separate list due to its length
    • Removed individual pages for families into one long list, families accessed from left menu via anchor redirects within page

Page created: February 02, 2002; Last edited: November 04, 2013 (EB)