Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

DOLICHIPODIDAE Latreille, 1809 - Long-legged flies

Often small, pretty metallic green or bronze flies that frequently are seen alighting upon plants, this is a large family of over 6,000 species in 200 genera. The great majority are terrestrial, but some groups are found in the margins of water bodies or in saturated soils nearby, often in large numbers. Aquatic representatives have larvae occuring in still or slow-moving waters, usually near the water margins in soft mud or decaying organic matter. Larvae and adults are mostly predacious on soft-bodies invertebrates.

Records are based on a survey of material at UMMZ-Insect Division collection, and literature records including Foote et al. (1965) and Pollet et al. (2004)

Amblypsilopus Bigot, 1888a: xxiv (Subfamily Sciapodinae)
Perhaps found around lake margins.
Amblypsilopus fuscinervis (Van Duzee, 1926c: 56 as Psilopus) - ON (Point Pelee; Pillot et al. 2004: 68)
Amblypsilopus scintillans (Loew, 1861: 96 as Psilopus) - UMMZ-Insect Division (as Sciapus scintillans)
(syn.) Psilopus angustatus Enderlein, 1912: 405
(syn.) Sciapus fulgidus Parent, 1929: 242
(syn.) Sciapus dubiosus Van Duzee, 1932b: 8
Amplypsilopus unifasciatus (Say, 1823: 85 as Dolichopus) - UMMZ-Insect Division (Sciapus unifasciatus); Point Pelee (Pillot et al. 2004: 68)
(syn.) Psilopus sayi Wiedemann, 1830: 219
(syn.) Psilopus ungulivena Walker, 1857-1860: 149
(syn.) Psilopus amabilis Parent, 1929: 242
(syn.) Chrysosoma plumosa Van Duzee, 1932a: 1

Argyra Macquart, 1834: 456 (Subfamily Diaphorinae)
Adults are hygrophilous and are frequently encountered on moist soil and herbaceous foliage near water, especially running water. Males, in particular, have silver pruinosity and resemble silvery flashes when in flight.
Argyra albicans Loew, 1861: 45 - UMMZ-Insect Division; NE-MN to QB, south to KS-GA (Pollet et al. 2004: 19)
(nom.nud.) Porphyrops quadriplagiatus Harris, 1835: 597
Argyra albicoxa Van Duzee, 1925a: 30 - IA-QB (Pollet et al. 2004: 19)
Argyra angustata Van Duzee, 1925a: 10 - WI, to ME (Pollet et al. 2004: 19)
Argyra bimaculata Van Duzee, 1925a: 19 - IL, widespread (Pollet et al. 2004: 19)
Argyra calceata Loew, 1861: 47 - IL, IN, OH (Pollet et al. 2004: 19)
Argyra calcitrans Loew, 1861: 46 - UMMZ-Insect Division; IN, ON, widespread central and east (Pollet et al. 2004: 19)
Argyra cingulata (Loew, 1861: 53 as Leucostola) - IN, ME-KS (Pollet et al. 2004: 19)
Argyra currani Van Duzee, 1925a: 24 - Foote et al. 1965, Pollet et al. 2004: 19
Argyra flavipes Van Duzee, 1925a: 34 - IN, to NY (Pollet et al. 2004: 19)
Argyra inaequalis Van Duzee, 1925a: 40 - IN (Lafayette), TN (may be found in sw LP)(Pollet et al. 2004: 19)
Argyra involuta Van Duzee, 1925a: 38 - IN, to SD (may be found in sw LP)(Pollet et al. 2004: 19)
Argyra medusae Gosseries, 1988: 305 - IN: 305
(syn.) Argyra abdominalis (Say, 1829-1830: 170 as Dolichopus)
Argyra minuta Loew, 1861: 46 - IN, ON (Pollet et al. 2004: 19)
Argyra nigricoxa Van Duzee 1925d: 26 - IN, OH, ON (Pollet et al. 2004: 19)
Argyra robusta Johnson, 1906: 59 - UMMZ-Insect Division; widespread (Pollet et al. 2004: 20)
Argyra sericata Van Duzee, 1925a: 28 - IN, ME-IA (Pollet et al. 2004: 20)
Argyra thoracica Van Duzee, 1925a: 23 - IN, ON, TN-NY (Pollet et al. 2004: 20)
Argyra velutina Van Duzee, 1925a: 20 - Foote et al. 1965; (Pollet et al. 2004: 20)

Asyndetus Loew 1869a: 25 (Subfamily Diaphorinae)
Species primarily coastal, but a few are found around inland lakes.
Asyndetus ammophilus Loew 1869a: 34 - IL, IN, RI-IA (Pollet et al. 2004: 20)
Asyndetus appendiculatus Loew 1869a: 36 - Foote et al. 1965; Pollet et al. 2004: 20
Asyndetus latus Van Duzee 1916b: 91 - UMMZ-Insect Division, but Foote et al. 1965 and Pollet et al. 2004: 21 note MI record is questionable
Asyndetus syntormoides Wheeler 1899: 32 - MA-KS Pollet et al. 2004: 21

Calyxochaetus Bigot, 1888a: xxiv (Subfamily Sympycninae)
Adults are common in shady areas and frequently hover over moist rocks and falling water (Pollet et al. 2004).
Calyxochaetus frontalis (Loew, 1861: 67 as Sympycnus) - UMMZ-Insect Division; IA-east (Pollet et al. 2004: 74)
Calyxochaetus nodatus (Loew, 1862: 215 as Sympycnus) - UMMZ-Insect Division; IL, IN, MN, OH, ON, WI (Pollet et al. 2004: 74)
(syn.) Nothosympycnus abbreviatus Van Duzee, 1917: 341

Campsicnemus Haliday, 1851: 357 (Subfamily Sympycninae)
Species are small, found on moist sand or soil and on low herbaceous foliage near the margins of streams and lakes. Adults sometimes run about on the surface of still water.
Campsicnemus americanus Van Duzee, 1924b: 3 - Foote et al. 1965
Campsicnemus degener Wheeler, 1899: 58 - widespread north
Campsicnemus hirtipes Loew, 1861: 68 - UMMZ-Insect Division
(syn.) Campsicnemus latipes Curran, 1934: 218 [lapsis]


Chrysotus Meigen, 1824: 40 (Subfamily Diaphorinae)
Adults usually are found in sunny spots, but some prefer wet, sandy banks of ponds and rivers.
Chrysotus aldrichi (Van Duzee, 1915: 185 as Diaphorus) - record in Pollet et al. 2004: 21 is noted as questionable
Chrysotus argentatus Van Duzee, 1924a: 41 - Foote et al. 1965, Pollet et al. 2004: 21
Chrysotus bellus Van Duzee, 1924a: 39 - Foote et al. 1965, Pollet et al. 2004: 21
Chrysotus canadensis Van Duzee, 1924a: 19 - UMMZ-Insect Division; Pollet et al. 2004: 22
Chrysotus choricus Wheeler, 1890: 357 - IL, IN, ON, WI (Pollet et al. 2004: 22)
(syn.) Chrysotus ciliatus Malloch, 1914: 236
Chrysotus convergens Van Duzee, 1924a: 17 - Foote et al. 1965; Pollet et al. 2004: 22
Chrysotus cornutus Loew, 1862: 214 - IL, IA-NY (Pollet et al. 2004: 22)
Chrysotus costalis Loew, 1861: 64 - UMMZ-Insect Division; Pollet et al. 2004: 22
Chrysotus discolor Loew, 1861: 65 - UMMZ-Insect Division; widespread (Pollet et al. 2004: 22)
(syn.) Chrysotus cobaltinus Van Duzee, 1924a: 38
Chrysotus distinctus Van Duzee, 1924a: 23 - UMMZ-Insect Division; Pollet et al. 2004: 22
Chrysotus dividuus Van Duzee, 1924a: 12 - Foote et al. 1965; Pollet et al. 2004: 22
Chrysotus hirtipes Van Duzee, 1924a: 22 - UMMZ-Insect Division; Pollet et al. 2004: 23
Chrysotus johnsoni Van Duzee, 1924a: 21 - Foote et al. 1965; Pollet et al. 2004: 23
Chrysotus leucostoma (Loew, 1861: 58 as Diaphorus) - UMMZ-Insect Division; widespread (Pollet et al. 2004: 23)
(syn.) Diaphorus infuscatus (Van Duzee, 1915: 179)
(syn.) Diaphorus vittatus (Van Duzee, 1915: 175)
Chrysotus megalocerus Meuffels & Grootaert, 1999: 289 - IA-NY (Pollet et al. 2004: 23)
(syn.) Chrysotus magnicornis Van Duzee 1924b: 13, preocc. by C. magnicornis Zetterstedt, 1843)
Chrysotus millardi Meuffels & Grootaert, 1999: 289 - Pollet et al. 2004: 23
(syn.) Chrysotus annulatus Van Duzee, 1924a: 27
Chrysotus obliquus Loew, 1861: 63 - UMMZ-Insect Division; widespread (Pollet et al. 2004: 23)
Chrysotus pallipes Loew, 1861: 66 - UMMZ-Insect Division; IL, NE-CT (Pollet et al. 2004: 23-24)
Chrysotus palpiger (Wheeler, 1890: 360 as Diaphorus) - UMMZ-Insect Division; widespread (Pollet et al. 2004: 24)
(syn.) Chrysotus spinifer Malloch, 1914: 238
Chrysotus philtrum Melander, 1903: 72 - UMMZ-Insect Division; Pollet et al. 2004: 24
Chrysotus picticornis Loew, 1862: 214 - UMMZ-Insect Division; widespread (Pollet et al. 2004: 24)
Chrysotus sodalis (Loew, 1861: 58 as Diaphorus) - OH, WI, IA-NH south (Pollet et al. 2004: 24)
Chrysotus spectabilis (Loew, 1861: 57 as Diaphorus) - widespread (Pollet et al. 2004: 24)
(syn.) Diaphorus exunguis Thomson, 1869: 506
(syn.) Diaphorus approximatus Aldrich, 1896: 321
Chrysotus subcostatus Loew, 1864b: 181- Foote et al. 1965; Pollet et al. 2004: 24
(syn.) Chrysotus pratincola Wheeler, 1890
Chrysotus tibialis Van Duzee, 1924a: 16 - UMMZ-Insect Division; Pollet et al. 2004: 24
Chrysotus variabilis (Van Duzee, 1915: 191 as Diaphorus) - UMMZ-Insect Division; IL, MN, NE-NY south (Pollet et al. 2004: 24-25)
Chrysotus vulgaris Van Duzee, 1924a: 15 - Foote et al. 1965; Pollet et al. 2004: 25
Chrysotus wisconsinensis Wheeler, 1890: 356 - UMMZ-Insect Division; WI-QB (Pollet et al. 2004: 25)

Diaphorus Meigen, 1824: 32 (Subfamily Diaphorinae)
This genus includes both hygrophilous and xerophilous species which are known from a variety of different habitats (heathland, river banks, humid forests).
Diaphorus gibbosus Van Duzee, 1915: 173 - UMMZ-Insect Division; widespread (Pollet et al. 2004: 25)
Diaphorus pseudopacus Robinson, 1964: 141 - IN, ON, WI, widespread (Pollet et al. 2004: 26)

Dolichopus Latreille 1796: 159 (Subfamily Dolichopodinae)
Prefers humid habitats, marshes, margins of lakes and streams.
Dolichopus abruptus Aldrich, 1922: 14 - UMMZ-Insect Division; Pollet et al. 2004: 27
Dolichopus absonus Van Duzee, 1921b: 294 - UMMZ-Insect Division; Pollet et al. 2004: 27
Dolichopus acuminatus Loew, 1861: 12 - UMMZ-Insect Division; Pollet et al. 2004: 27
Dolichopus adultus Van Duzee, 1921b: 98 - UMMZ-Insect Division; Pollet et al. 2004: 27-28
Dolichopus aethiops Van Duzee, 1921b: 296 - IN, Lafayette (Pollet et al. 2004: 28)
Dolichopus affinis Walker, 1848-1849: 659 - UMMZ-Insect Division; Pollet et al. 2004: 28
(syn.) Dolichopus splendidulus Loew, 1864a: 91
Dolichopus ainsliei Van Duzee, 1921b: 272 - MN, WI, east to NY (Pollet et al. 2004: 28)
Dolichopus alacer Van Duzee, 1921b: 86 - IN, Lafayette, ON (Pollet et al. 2004: 28)
Dolichopus albiciliatus Loew, 1862: 211 - UMMZ-Insect Division: Pollet et al. 2004: 28
Dolichopus albicoxa Aldrich, 1893: 10 - UMMZ-Insect Division; Pollet et al. 2004: 28
Dolichopus amphericus Melander & Brues, 1900: 146 - WI, northern (Pollet et al. 2004: 28)
Dolichopus amplipennis Van Duzee, 1921b: 197 - UMMZ-Insect Division; Pollet et al. 2004: 28
Dolichopus angustatus Aldrich, 1893: 15 - MN, eastward (Pollet et al. 2004: 28)
Dolichopus apheles Melander & Brues, 1900: 144 - Foote et al. 1965; Pollet et al. 2004: 28
Dolichopus barbicauda Van Duzee, 1921b: 76 - UMMZ-Insect Division; Pollet et al. 2004: 28-29
Dolichopus bifractus Loew, 1861: 19 - UMMZ-Insect Division; Pollet et al. 2004: 29
Dolichopus brevimanus Loew, 1861: 14 - WI, widespread (Pollet et al. 2004: 29)
Dolichopus brevipennis Meigen, 1824: 89 - widespread northern, expect in boreal UP (Pollet et al. 2004: 29)
Dolichopus bruesi Van Duzee, 1921 - WI, westward (UP?) (Pollet et al. 2004: 29)
(syn.) Hygroceleuthes propinquus (Melander & Brues, 1900) (preocc. by Dolichopus propinquus Zetterstedt, 1852)
Dolichopus burnesi Van Duzee, 1921b: 64 - UMMZ-Insect Division; Pollet et al. 2004: 29
Dolichopus calainus Melander & Brues, 1900: 138 - IL (Pollet et al. 2004: 29)
Dolichopus calcaratus Aldrich, 1893: 8 - UMMZ-Insect Division; ON, IA-QB (Pollet et al. 2004: 29)
Dolichopus canadensis Van Duzee, 1921b: 141 - UMMZ-Insect Division; Pollet et al. 2004: 29
Dolichopus coercens Walker, 1848-1849: 661 - UMMZ-Insect Division; IL, ON, WI (Pollet et al. 2004: 29-30)
(syn.) Dolichopus conterminus Walker, 1848-1849: 664
(syn.) Dolichopus batillifer Loew, 1861: 15
Dolichopus coloradensis Aldrich, 1893: 23 - CO-QB, northern (Pollet et al. 2004: 30)
Dolichopus comatus Loew, 1861: 23 - UMMZ-Insect Division; IL, IN, OH, WI (Pollet et al. 2004: 30)
Dolichopus contiguus Walker, 1848-1849: 663 - UMMZ-Insect Division; Pollet et al. 2004: 30
(syn.) Dolichopus splendilus Loew, 1861: 14
Dolichopus coquilletti Aldrich, 1893: 19 - widespread (Pollet et al. 2004: 30)
Dolichopus correus Steyskal, 1959: 1 - UMMZ-Insect Division; Pollet et al. 2004: 30
Dolichopus cuprinus Wiedemann, 1830: 230 - UMMZ-Insect Division; Pollet et al. 2004: 30
Dolichopus dakotensis Aldrich, 1893: 11 - UMMZ-Insect Division; IL, MB-SD to QB (Pollet et al. 2004: 30)
Dolichopus dasypodus Coquillett, 1910: 42 - UMMZ-Insect Division; Pollet et al. 2004: 30
Dolichopus decorus Van Duzee, 1921b: 153 - UMMZ-Insect Division; Pollet et al. 2004: 30
Dolichopus defectus Van Duzee, 1921b: 143 - ON, MB-QB (Pollet et al. 2004: 30)
Dolichopus detersus Loew, 1866: 44 - UMMZ-Insect Division; AK-QB (Pollet et al. 2004: 31)
Dolichopus discolor Van Duzee, 1921b: 116 - UMMZ-Insect Division; IL, ON (Pollet et al. 2004: 31)
Dolichopus dolosus Parent, 1934b: 267 - WI (Pollet et al. 2004: 31)
Dolichopus domesticus Van Duzee, 1921b: 246 - UMMZ-Insect Division; Pollet et al. 2004: 31
Dolichopus enigma Melander & Brues, 1900: 139 - Foote et al. 1965; Pollet et al. 2004: 31
Dolichopus eudactylus Loew, 1861: 16 - UMMZ-Insect Division; Pollet et al. 2004: 31
Dolichopus factivittatus Harmston, 1966: 10 - IN, Terre Haute (Pollet et al. 2004: 31)
Dolichopus finitus Walker, 1848-1849: 662 - UMMZ-Insect Division; Pollet et al. 2004: 31
(nom.nud.) Dolichopus atricornis Harris, 1835: 597
(syn.) Dolichopus scoparius Loew, 1864b: 70
Dolichopus flagellitenens Wheeler, 1890: 339 - UMMZ-Insect Division; WI, SD-NH (Pollet et al. 2004: 31)
Dolichopus flavicoxa Van Duzee, 1921b: 188 - UMMZ-Insect Division; WI, BC-NY (Pollet et al. 2004: 31)
Dolichopus fulvipes Loew, 1862: 212 - UMMZ-Insect Division; IL, AB-QB (Pollet et al. 2004: 32)
Dolichopus gladius Van Duzee, 1921b: 136 - UMMZ-Insect Division; Pollet et al. 2004: 32
Dolichopus gratus Loew, 1861: 11 - UMMZ-Insect Division; Pollet et al. 2004: 32
Dolichopus imperfectus Van Duzee, 1921b: 240 - IL, Calumet, expect in sw LP (Pollet et al. 2004: 32)
Dolichopus incisuralis Loew, 1861: 25 - UMMZ-Insect Division; IN, ON, NT-NH south (Pollet et al. 2004: 32)
(syn.) Dolichopus platyprosopus Loew, 1866: 44
Dolichopus incongruus Wheeler, 1890: 338 - WI, ON, south to TN (Pollet et al. 2004: 32)
Dolichopus indigenus Van Duzee, 1921b: 139 - MN, widespread northern (Pollet et al. 2004: 32)
Dolichopus intentus Melander & Brues, 1900: 137 - IL, Chicago (Pollet et al. 2004: 32)
Dolichopus kleini Curran, in Van Duzee & Curran, 1934: 25 - MN, OH (Pollet et al. 2004: 33)
Dolichopus laciniatus Coquillet, 1910: 42 - Foote et al. 1965; ON, IA-NY (Pollet et al. 2004: 33)
Dolichopus laticornis Loew, 1861: 12 - UMMZ-Insect Division; Pollet et al. 2004: 33
Dolichopus latipes (Loew, 1861: 5 as Hygroceleuthus) - UMMZ-Insect Division; Pollet et al. 2004: 33
Dolichopus lobatus Loew, 1861: 24 - UMMZ-Insect Division; Pollet et al. 2004: 33
Dolichopus longimanus Loew, 1861: 14 - UMMZ-Insect Division; BC-NH (Pollet et al. 2004: 33)
Dolichopus longipennis Loew, 1861: 21 - UMMZ-Insect Division; Pollet et al. 2004: 33
Dolichopus luteipennis Loew, 1861: 18 - UMMZ-Insect Division; Pollet et al. 2004: 33
Dolichopus melanocerus Loew, 1864a: 93 - Foote et al. 1965; Pollet et al. 2004: 34
Dolichopus multisetosus Van Duzee, 1921b: 49 - western, but questionable record in ME (Pollet et al. 2004: 34), perhaps widespread?
Dolichopus nigricornis Meigen, 1824: 82 - UMMZ-Insect Division; AK-QB (Pollet et al. 2004: 34)
(syn.) Dolichopus discifer Stannius, 1831: 57
(syn.) Dolichopus tanypus Loew, 1861: 24
Dolichopus nodipennis Van Duzee, 1921b: 102 - UMMZ-Insect Division; ON, WI (Pollet et al. 2004: 34)
Dolichopus nubifer Van Duzee, 1921b: 75 - UMMZ-Insect Division; Pollet et al. 2004: 34
Dolichopus obcordatus Aldrich, 1893: 14 - UMMZ-Insect Division; Pollet et al. 2004: 34
Dolichopus occidentalis Aldrich, 1893: 19 - western, but questionable record in PEI, perhaps widespread? (Pollet et al. 2004: 34)
Dolichopus omnivagus Van Duzee, 1921b: 216 - UMMZ-Insect Division; AK-ME, s. to IA (Pollet et al. 2004: 35)
Dolichopus orichalceus Gosseries, 1988: 304 - Pollet et al. 2004: 35
(syn.) Dolichopus vittatus Loew, 1861: 20 (preocc. by D. vittatus Wiedemann, now Chrysosoma)
Dolichopus ovatus Loew, 1861: 13 - UMMZ-Insect Division; Pollet et al. 2004: 35
Dolichopus pachycnemus Loew, 1861: 13 - UMMZ-Insect Division; IL, MB-QB (Pollet et al. 2004: 35)
Dolichopus palaestricus Loew, 1864a: 92 - UMMZ-Insect Division; Pollet et al. 2004: 35
Dolichopus parvicornis Van Duzee, 1921b: 231 - MN, ON (Pollet et al. 2004: 35)
Dolichopus penicillatus Van Duzee, 1921b: 227 - widespread (Pollet et al. 2004: 35)
(syn.) Dolichopus ciliatus Aldrich, 1893: 25
Dolichopus plumipes (Scopoli, 1763: 224 as Musca) - UMMZ-Insect Division; Pollet et al. 2004: 35-36
(syn.) Dolichopus pennitaris Fallén, 1823: 11
(syn.) Dolichopus ciliatus Walker, 1848-1849: 661
(syn.) Dolichopus sequax Walker, 1848-1849: 666
(syn.) Dolichopus nigriapicalis Van Duzee, 1930a: 125
Dolichopus porphyrops Van Duzee, 1921b: 169 - UMMZ-Insect Division; Pollet et al. 2004: 36
Dolichopus praeustus Loew, 1862: 212 - IL, IN, to CN (Pollet et al. 2004: 36)
Dolichopus pulchrimanus (Bigot, 1888b: xxx as Spathichira) - UMMZ-Insect Division; Pollet et al. 2004: 36
(syn.) Dolichopus willistonii Aldrich, 1893: 22
Dolichopus ramifer Loew, 1861: 19 - UMMZ-Insect Division; Pollet et al. 2004: 36
Dolichopus reflectus Aldrich, 1893: 12 - UMMZ-Insect Division; Pollet et al. 2004: 36
Dolichopus remipes Wahlberg, 1839: 13 - UMMZ-Insect Division; ON, WI, widespread north (Pollet et al. 2004: 36)
Dolichopus remus Van Duzee, 1921b: 96 - UMMZ-Insect Division; IL, MN, OH, ON (Pollet et al. 2004: 36)
Dolichopus renidescens Melander & Brues, 1900: 143 - widespread west to MN, but questionable record from ME (Pollet et al. 2004: 36)
Dolichopus retinens Van Duzee, 1921b: 90 - IN, Lafayette, ON, widespread (Pollet et al. 2004: 36)
Dolichopus ruficornis Loew, 1861: 21 - UMMZ-Insect Division; Pollet et al. 2004: 36
Dolichopus sarotes Loew, 1866: 44 - UMMZ-Insect Division; IL, IN, ON, WI (Pollet et al. 2004: 36)
Dolichopus scapularis Loew, 1861: 22 - Pollet et al. 2004: 36-37
(syn.) Dolichopus alberecrus Gunther, 1979: 1
Dolichopus sedulus Van Duzee, 1921b: 93 - IN, north (Pollet et al. 2004: 37)
Dolichopus serratus Van Duzee, 1921b: 155 - Foote et al. 1965; Pollet et al. 2004: 37
Dolichopus setifer Loew, 1861: 12 - UMMZ-Insect Division; Pollet et al. 2004: 37
Dolichopus setosus Loew, 1862: 213 - eastern, ON, IA, questionable record from BC (Pollet et al. 2004: 37)
Dolichopus sexarticulatus Loew, 1864b: 62 - Foote et al. 1965; Pollet et al. 2004: 37
Dolichopus sicarius Van Duzee, 1921b: 261 - Foote et al. 1965; Pollet et al. 2004: 37
Dolichopus sincerus Melander, 1900: 136 - UMMZ-Insect Division; ON, WI, OR-ME (Pollet et al. 2004: 37)
Dolichopus socius Loew, 1862: 211 - UMMZ-Insect Division; IL, OR-QB, NS (Pollet et al. 2004: 37)
Dolichopus stenhammari Zetterstedt, 1843: 521 - widespread north, expect in UP (Pollet et al. 2004: 37)
(syn.) Dolichopus annulipes Zetterstedt, 1838: 710 (preocc. in Dolichopus by Meigen, 1824, orig. Porphyrops, now Sympycnus)
Dolichopus tener Loew, 1861: 17 - UMMZ-Insect Division; IL, ON, WI (Pollet et al. 2004: 38)
Dolichopus terminalis Loew, 1866: 43 - UMMZ-Insect Division; ON, WI (Pollet et al. 2004: 38)
(syn.) Dolichopus germanus Wheeler, 1890: 341 [preocc. by D. germanus Wiedemann, 1817, now Hercostomus]
Dolichopus terminatus Walker, 1848-1849: 665 - widespread? (Pollet et al. 2004: 38)
Dolichopus tetricus Loew, 1864b: 33 - widespread boreal? (Pollet et al. 2004: 38)
Dolichopus tonsus Loew, 1861: 16 - UMMZ-Insect Division; Pollet et al. 2004: 38
Dolichopus umbrosus Van Duzee, 1921b: 100 - ON, WI (Pollet et al. 2004: 38)
Dolichopus variabilis Loew, 1861: 17 - UMMZ-Insect Division; Pollet et al. 2004: 38
(syn.) Dolichopus gracilis Aldrich, 1893: 15
(syn.) Dolichopus annaclareii Gunther, 1979: 1
Dolichopus versutus Van Duzee, 1921b: 253 - UMMZ-Insect Division; IN, ON, KS-QB (Pollet et al. 2004: 38)
Dolichopus vigilans Aldrich, 1893: 13 - IN, OH, ON, southward to KS (Pollet et al. 2004: 38)
Dolichopus wheeleri (Melander & Brues, 1900: 126 as Hygroceleuthus) - MB-NS (Pollet et al. 2004: 39)
Dolichopus xanthocnemus Loew, 1864b: 31- ON, widespread north (Pollet et al. 2004: 39)

Gymnopternus Loew, 1857a: 10 (Subfamily Dolichopodinae)
Most diverse in humid forests. Variously treated as a subgenus or synonym of Hercostomus.
Gymnopternus anarmostus (Melander, 1900: 139 as Hercostomus) - IL, Chicago (Pollet et al. 2004: 39)
Gymnopterus annulatus Van Duzee, 1926c: 58 - Pollet et al. 2004: 39
Gymnopterus annulipes Robinson, 1964: 161 - Pollet et al. 2004: 39
Gymnopterus barbatulus Loew, 1861: 29 - UMMZ-Insect Division; Pollet et al. 2004: 39
Gymnopterus crassicauda Loew, 1861: 35 - UMMZ-Insect Division; OH, OH, IA-QB (Pollet et al. 2004: 39)
Gymnopterus debilis Loew, 1861: 35 - Pollet et al. 2004: 40
Gymnopternus difficilis Loew, 1861: 33 - ON, NS-IA (Pollet et al. 2004: 40)
Gymnopternus exilis Loew, 1861: 30 - IN-NH (Pollet et al. 2004: 40)
Gymnopternus flavus Loew, 1861: 28 - ON, IN, QB-GA (Pollet et al. 2004: 40)
Gymnopternus frequens Loew, 1861: 32 - UMMZ-Insect Division; Pollet et al. 2004: 40
Gymnopternus humilis Loew, 1864b: 336 - UMMZ-Insect Division; Pollet et al. 2004: 40
Gymnopternus lividifrons Van Duzee, 1926c: 58 - Pollet et al. 2004: 40
Gymnopternus lunifer Loew, 1861: 32 - IL, east to QB (Pollet et al. 2004: 40)
(syn.) Gymnopternus exiguus Loew, 1864b: 337
Gymnopternus nigribarbus Loew, 1861: 33 - UMMZ-Insect Division; Pollet et al. 2004: 40
Gymnopternus opacus Loew, 1861: 34 - IA-QB (Pollet et al. 2004: 41)
Gymnopternus politus Loew, 1861: 34 - Foote et al. 1965; Pollet et al. 2004: 41
Gymnopternus purpuratus (Van Duzee, 1925b: 185 as Hercostomus) - Pollet et al. 2004: 41
Gymnopternus scotias Loew, 1861: 29 - ON, NS-IA-MB (Pollet et al. 2004: 41)
Gymnopternus spectabilis Loew, 1861: 30 - UMMZ-Insect Division (also as Diaphorus spectabilis Loew); ON, WI, east (Pollet et al. 2004: 41)
(syn.) Gymnopternus chalcochrus Loew, 1861b: 335
Gymnopternus violaceus (Van Duzee, 1921a: 123 as Proarchus) - Pollet et al. 2004: 41

Hercostomus Loew 1857a: 9 (Subfamily Dolichopodinae)
Representatives are found in a wide variety of habitats, ranging from coastal dunes to humid forests and springs.
Hercostomus chetifer Walker 1848-1849: 653 - Pollet et al. 2004: 42
(syn.) Dolichopus alutifer Haliday, 1851: 179
(syn.) Paraclius ornatus Van Duzee, 1921a: 128
(syn.) Hercostomus dreisbachi Harmston & Knowlton, 1945: 80
Hercostomus costalis Van Duzee, 1923b: 65 - Foote et al. 1965; Pollet et al. 2004: 42
Hercostomus flutatus Harmston & Knowlton, 1945: 79 - Harmston & Knowlton 1945: 79; Pollet et al. 2004: 42
Hercostomus tibialis (Van Duzee, 1913: 55 as Neurigona) - IL, IN, ON (Pollet et al. 2004: 43)
(syn.) Neurigona dorsalis Van Duzee, 1914: 434
(syn.) Hercostomus indianus Harmston, 1952: 285
Hercostomus unicolor Loew, 1864b: 117 - Pollet et al. 2004: 43
(syn.) Gymnopternus poenitens Wheeler, 1890: 355
(syn.) Hercostomus ornaticauda Van Duzee, 1933a: 16

Hydatostega Philippi 1865: 779 (Subfamily Hydrophorinae)
Adults are generally associated with standing water and are adept at skating on the surface.
Hydatostega viridiflos viridiflos (Walker, 1852-1853: 212 as Medeterus) - QB/NY-MB (Pollet et al. 2004: 49)
(syn.) Hydrophorus intentus Aldrich, 1911: 51
(syn.) Hydrophorus indentus, error by Foote et al. in listing H. intentus Aldrich, 1911
(syn.) Hydrophorus browni Curran, 1930: 73

Hydrophorus Fallén, 1823: 2 (Subfamily Hydrophorinae)
Adults of some species are seen skating over quiet surface of large, stagnant bodies of water, and others are found on the shores of large rivers and streams.
Hydrophorus aestuum Loew, 1869: 36 - Pollet et al. 2004: 49
Hydrophorus agalma Wheeler, 1899: 66 - UMMZ-Insect Division; Pollet et al. 2004: 49
Hydrophorus alboflorens Walker, 1848-1849: 656 - UMMZ-Insect Division; Pollet et al. 2004: 49
(syn.) Hydrophorus pirata Loew, 1861: 71
Hydrophorus algens Wheeler, 1899: 63 - widespread northern, expect in UP (Pollet et al. 2004: 49)
(syn.) Dydrophorus hirtipes Van Duzee, 1933b: 17
Hydrophorus altivagus Aldrich, 1911: 67 - MN, ON, widespread northern (Pollet et al. 2004: 49)
(syn.) Hydrophorus wahlgreni Frey, 1915: 4, 57, 63
(syn.) Hydrophorus lividipes Van Duzee, 1926b: 50
Hydrophorus amplectens Aldrich, 1911: 67 - Foote et al. 1965; Pollet et al. 2004: 50
Hydrophorus angustifacies Hurley, 1985: 29 - Pollet et al. 2004: 50
Hydrophorus aquatilis Aldrich, 1922: 17 - widespread northern
(syn.) Hydrophorus criddlei Van Duzee, 1925b: 181
(syn.) Hydrophorus argentifacies Van Duzee, 1926b: 51
Hydrophorus chrysologus (Walker, 1848-1849: 655 as Medeterus) - UMMZ-Insect Division; Pollet et al. 2004: 50
(syn.) Hydrophorus glaber not Walker, misid. by Wheeler (1899); error for H. glaber by Van Duzee (1923)
(syn.) Hydrophorus algens not Wheeler, misid. by Snodgrass (1904)
(syn.) Hydrophorus alter Parent, 1934a: 19
Hydrophorus dreisbachi Harmston & Knowlton, 1963: 231 - Harmston & Knowlton 1963: 231; Pollet et al. 2004: 50
Hydrophorus flavihirtus Van Duzee, 1923a: 259 - Pollet et al. 2004: 50
Hydrophorus flavipennis Van Duzee, 1926b: 48 - Foote et al. 1965; Pollet et al. 2004: 51
Hydrophorus parvus Loew 1862: 214 - Foote et al. 1965; Pollet et al. 2004: 51
Hydrophorus philombrius Wheeler, 1890: 378 - UMMZ-Insect Division; Pollet et al. 2004: 51
Hydrophorus sodalis Wheeler, 1899: 68 - Foote et al. 1965; Pollet et al. 2004: 51-52

Lamprochromus Mik, 1878: 7 (Subfamily Sympycninae)
In the Palearctic, L. bifasciatus and L. strobli are almost entirely restricted to reed marshes (Pollet, 2000). The habits of the Nearctic species are unknown.
Lamprochromus canadensis (Van Duzee, 1917: 339 as Sympycnus) - ON, widespread northern (Pollet et al. 2004: 76)

Liancalus Loew 1857a: 22 (Subfamily Hydrophorinae)
Species are hygrophilus, often found in association with vertical seepages. Adults of L. similis are found in moist shaded areas in association with drainage culverts, freshet seeps, and waterfalls. Larvae are known from algal mats (Corpus, 1986).
Liancalus genualis Loew, 1861: 70 - Pollet et al. 2004: 52

Nematoproctus Loew, 1857a: 40 (Subfamily Rhaphiinae)
Adults are hydrophilous and occur on river banks and in humid deciduous forests, where they can be found on herbaceous foliage.
Nematoproctus metallicus Van Duzee, 1930c: 168 - UMMZ-Insect Division; Pollet et al. 2004: 64
Nematoproctus venustus Melander, 1900: 142 - eastern, but a questionable record from IA (Pollet et al. 2004: 64)

Nepalomyia Hollis, 1964: 110 (Subfamily Peloropeodinae)
Adults seem to be hygrophilous and prefer shaded habitats with seepages on moss-covered rocks (Runyon and Hurley 2003).
Nepalomyia sombrea (Harmston & Knowlton, 1945: 77 as Neurigona) - Foote et al. 1965, Midland Co., June 1941, RR Dreisbach (Runyon & Hurley 2003); Pollet et al. 2004: 63

Paraclius Loew, 1864b: 97 (Subfamily Dolichopodinae)
Most species of this genus are maritime, but several species are found in our area, near margins of lakes and streams.
Paraclius ovatus Van Duzee, 1914: 436 - Foote et al. 1965; Pollet et al. 2004: 43
Paraclius pumilio Loew, 1872: 90 - Foote et al. 1965 ; Pollet et al. 2004: 44

Pelastoneurus Loew, 1861: 36 (Subfamily Dolichopodinae)
Species are found in association with freshwater seepages and mudflats, and on wet soil or on low herbaceous vegetation near water.
Pelastoneurus abbreviatus Loew, 1864a; 94 - IL, ON (Pollet et al. 2004: 44)
Pelastoneurus kansensis (Aldrich, 1894: 153 as Metapelastoneurus) - UMMZ-Insect Division; Pollet et al. 2004: 44
Pelastoneurus laetus Loew, 1861: 38 - Foote et al. 1965; Pollet et al. 2004: 44-45
Pelastoneurus lamellatus Loew, 1864a: 95 - UMMZ-Insect Division; NY-IA south (Pollet et al. 2004: 45)
(syn.) Pelastoneurus cristatus Van Duzee, 1924a: 105
Pelastoneurus lugubris Loew, 1861: 38 - Foote et al. 1965; Pollet et al. 2004: 45
Pelastoneurus neglectus Wheeler, 1899: 12 - UMMZ-Insect Division; Pollet et al. 2004: 45
Pelastoneurus proximus Aldrich, 1904: 278 - Foote et al. 1965
Pelastoneurus umbripictus Becker, 1922: 73 - Foote et al. 1965: Pollet et al. 2004: 45
Pelastoneurus wheeleri Melander, 1900: 140 - Foote et al. 1965; Pollet et al. 2004: 45

Peloropeodes Wheeler, 1890: 373 (Subfamily Peloropeodinae)
Adults are found on low herbaceous vegetation, moist soil, and on rock surfaces near, or on, stream banks.
Peloropeodes aldrichi Robinson, 1970: 26, new name for Kophosoma acuticornis (Van Duzee, 1926) - Foote et al. 1965; Pollet et al. 2004: 63
(syn.) Kophosoma acuticornis Van Duzee, 1926a: 44, preocc. by Anomalopyga acuticornis Oldenberg, 1916, now Peloropeodes
Peloropeodes cornuta (Van Duzee, 1926a: 42 as Kophosoma) - Foote et al. 1965: Pollet et al. 2004: 63
Peloropeodes salax Wheeler, 1890: 374 - WI, Milwaukee, east (Pollet et al. 2004: 63)
(syn.) Peloropeodes flavipes Van Duzee, 1914: 437

Plagioneurus Loew, 1857b: 43 (Subfamily Plagioneuriinae)
Plagioneurus univittatus Loew, 1857b: 43 - UMMZ-Insect Division; Pollet et al. 2004: 64

Rhaphium Meigen, 1803: 272 (Subfamily Rhaphiinae)
Most species emerge early in the season and are found in moist habitats such as humid woodlands, river banks, and pond margins.
Rhaphium atkinsoni Curran, 1926: 257 - Foote et al. 1965; Pollet et al. 2004: 64
Rhaphium effilatum (Wheeler, 1899: 34 as Porphyrops) - UMMZ-Insect Division; WI, widespread (Pollet et al. 2004: 65)
Rhaphium elegantulum (Meigen, 1824: 51 as Porphyrops) - widespread northern (Pollet et al. 2004: 65)
Rhaphium exile Curran, 1926: 257 - IN, Lafayette (Pollet et al. 2004: 65)
Rhaphium fascipes (Meigen, 1824: 54 as Porphyrops (orig.) - UMMZ-Insect Division; Pollet et al. 2004: 65
Rhaphium femineum (Van Duzee, 1922: 84 as Xiphandrium) - Foote et al. 1965; Pollet et al. 2004: 65
Rhaphium melampus Loew, 1861: 50 as Porphyrops) - UMMZ-Insect Division; Pollet et al. 2004: 66
Rhaphium punctitarse Curran, 1924: 135 - UMMZ-Insect Division; Pollet et al. 2004: 67
Rhaphium rotundiceps (Loew, 1861: 51 as Porphyrops) - UMMZ-Insect Division; Pollet et al. 2004: 67
Rhaphium signiferum (Osten Sacken, 1878: 242 as Porphyrops) - Foote et al. 1965; Pollet et al. 2004: 67
Rhaphium vanduzeei Curran, 1926: 253 - IN-Lafayette, ON, NE-QB (Pollet et al. 2004: 67)
Rhaphium wheeleri Van Duzee, 1932a: 9- WI, Milwaukee (Pollet et al. 2004: 67)


Scellus Loew, 1857a: 22 (Subfamily Hydrophorinae)
Adults have been collected in swampy, high-mountain meadows, pastures on the plains, and on the bark of trees (Hurley, 1995).
Scellus exustus (Walker, 1852-1853: 211 as Medeterus) - UMMZ-Insect Division; Pollet et al. 2004: 54
(syn.) Scellus venustus, error by Crampton (1942: 94)

Sympycnus Loew, 1857a: 42 (Subfamily Sympycninae)
Adults of most species are more common in deep shade and frequently hover over moist rocks and falling water (Foote et al. 1965). Many species have been transferred to Calyxochaetus.
Sympycnus aldrichi Van Duzee, 1930b: 39 - IN, Lafayette, expect in s. LP (Pollet et al. 2004: 77)
Sympycnus lineatus Loew, 1861: 67 - UMMZ-Insect Division; MN-QB (Pollet et al. 2004: 77)

Syntormon Loew, 1857a: 35 (Subfamily Sympycninae)
Most species are riparian and mainly occur on the banks of stagnant or running water bodies, and in open, well-lit, humid habitats such as marshlands and saltmarshes (Pollet et al. 2004).
Syntormon cinereiventre (Loew, 1861: 48 as Synarthrus) - UMMZ-Insect Division; IN, ON, WI (Pollet et al. 2004: 78)
Syntormon femoratum Van Duzee, 1925a: 280 - IN, Lafayette, expect in s. LP (Pollet et al. 2004: 78)
Syntormon palmare (Loew, 1864b: 135 as Synarthrus) - Foote et al. 1965; Pollet et al. 2004: 79
Syntormon simplicitarse Van Duzee, 1925a: 286 - IN. Lafayette (Pollet et al. 2004: 79)
Syntormon tricoloripes Curran, 1923: 209 - Foote et al. 1965; Pollet et al. 2004: 79

Tachytrechus Stannius, 1831: 261 (Subfamily Dolichopodinae)
Species are hygrophilus and riparian, and mainly occur on open, well-lit, wet, sandy soils or rocky substrates near, or in, open water. Larval and pupal stages have been discovered along temporal pools (Dyte, 1959) and in freshwater seeps and mud flats (Corpus, 1988). Most species are western, but a few are found in our area.
Tachytrechus binodatus Loew, 1866: 46 - UMMZ-Insect Division; Pollet et al. 2004: 46
Tachytrechus moechus Loew, 1861: 40 - UMMZ-Insect Division; IL, IN, ON (Pollet et al. 2004: 47)
Tachytrechus vorax Loew, 1861: 41 - UMMZ-Insect Division ; Pollet et al. 2004: 47

Telmaturgus Mik, 1874: 349 (Subfamily Sympycninae)
Specimens of this species collected by Runyon (2012) were collected in Pennsylvania from deeply shaded sphagnum areas of a wooded swamp, most obtained by inserting a sweep net into dark areas underneath a boardwalk.
Telmaturgus parvus (Van Duzee, 1924a: 45 Chrysotus (orig.) - Foote et al. 1965; Pollet et al. 2004: 79

Teuchophorus Loew, 1857a: 44 (Subfamily Sympycninae)
Species of this genus are hygrophilous and, like Sympycnus, occur in rather shaded conditions where they can be found, sometimes in large numbers, on muddy soils (Pollet et al. 2004).
Teuchophorus clavigerellus Wheeler, 1899: 57 - UMMZ-Insect Division; Pollet et al. 2004: 79
Teuchophorus condylus Harmston & Knowlton, 1946: 571 - IN, ON (Pollet et al. 2004: 79)

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May 20, 2017

• Corrections to nomenclatures, organization and references.

Page created: April 25, 2006 - Last updated: May 20, 2017 (EB)