Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

Tipuloidea - Crane Flies

One of largest groups of Diptera, with more than 15,275 recognized species in 525 genera and subgenera (Oosterbroek, 2005). Traditionally considered the most basal of the Diptera and Nematocera (Wood and Borkent 1989, Michelsen 1996), evidence increasingly points to the Tipuloidea as a more derived nematoceran lineage (e.g. Oosterbroek and Courtney 1995). Literature often considers this groups as the Tipulidae (sensu lato), but more recently workers have elavated previous subfamilies to family level - Cylindrotomidae, Limoniidae, Pediciidae, and Tipulidae sensu stricto (Young 2008). However, an analysis of data from molecular sampling suggests that Limoniidae is para- and polyphyletic, and should be retained in Tipulidae (Petersen et al. 2010).

The majority of species are terrestrial, with immature life stages frequenting moist areas in soil, decayed organic substrates, or plant (usually woody) material. Other species live near the margins of lakes and streams, and are considered semiaquatic, with a few groups becoming adapted to strictly aquatic environments.

This list is based on works of Alexander and Stone et al. (1965), and should be considered only a beginning if not outdated taxonomic survey for the state. Determination of taxa considered aquatic or semi-aquatic is based on an interpretation of Rogers 1942 and Byers 1995. Based on literature records and museum specimens, there are 186 species in 73 genera of Tipuloidea associated with aquatic or wet habitats documented in Michigan, although this number is certainly on the low side of an actual inventory. Genera are denoted in green; species denoted in bold blue have been recorded in Michigan; species denoted in black bold are likely to occur in Michigan based on their known distribution with nearby state/provincial localities from which certified records have been published. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

CYLINDROTOMIDAE

Phalacrocera Schiner, 1863: 224 (Subfamily Cylindrotominae)
Phalacrocera neoxena Alexander, 1914: 603
Phalacrocera tipulina Osten Sacken, 1865: 241 - WI north and east


Triogma Schiner, 1863: 223 (Subfamily Cylindrotominae)
Triogma exsculpta Osten Sacken, 1865: 239

LIMONIIDAE
Antocha Osten Sacken, 1859: 219 (Tribe Limoniini)
Antocha obtusa Alexander, 1925b: 201 - Alexander, 1925
Antocha opalizans Osten Sacken, 1859: 220 - WI east
Antocha saxicola Osten Sacken, 1859: 219 - Kovalak 1978: 10


Arctoconopa Alexander, 1955: 17 (Tribe Eriopterini)
Arctoconopa painteri (Alexander, 1929c: 19 as Erioptera) - OH


Cryptolabis Osten Sacken, 1859: 224 (Tribe Eriopterini)
Cryptolabis (Phantolabis) lacustris (Alexander, 1938: 76 as Erioptera) - Alexander, 1938 (aquatic)


Dactylolabis Osten Sacken, 1859: 240 (Tribe Hexatomini)
[algal and other plant associations in dripping water on cliff faces or comparible associations (Alexander in Stone et al. 1985)]
Dactylolabis (Dactylolabis) cubitalis (Osten Sacken, 1869: 229 as Limnophila) - IN, OH, WI
Dactylolabis (Dactylolabis) montana (Osten Sacken, 1859: 240 as Limnophila) - IL east and northeast


Dicranota Zetterstedt, 1838: 851 (Tribe Pediciini)
Dicranota (Rhaphidolabina) flaveola (Osten Sacken, 1869: 288 as Rhaphidolabis)
Dicranota (Dicranota) currani Alexander, 1926b: 50 - MN, ON east
Dicranota (Dicranota) fumipennis Alexander, 1941: 358 - MN
Dicranota (Paradicranota) eucera Osten Sacken, 1869: 281 - MN east
Dicranota (Paradicranota) iowa Alexander, 1920a: 78 - MN south and east
Dicranota (Plectromyia) petiolata (Alexander, 1919b: 194 as Tricyphona) - CO northeast to QB
Dicranota (Rhaphidolabis) cayuga (Alexander, 1916a: 543 as Rhaphidolabis) - widespread
Dicranota (Rhaphidolabis) forceps (Alexander, 1924a: 63 as Rhaphidolabis)
Dicranota (Rhaphidolabis) rogersiana (Alexander, 1925b: 204 as Rhaphidolabis)
Dicranota (Rhaphidolabis) tenuipes (Osten Sacken, 1869: 287 as Rhaphidolabis)


Elliptera Schiner, 1863: 222 (Tribe Limoniini)
Elliptera illini Alexander, 1920c: 86 - IL


Erioptera Meigen, 1803: 262 (Tribe Eriopterini)
Erioptera (Trimicra) pilipes (Fabricius, 1787: 324 as Tipula) - NAm, Europe
(syn.) Trimicra anomala Osten Sacken, 1861: 290
Erioptera (Erioptera) chlorophylla Osten Sacken, 1859: 226 - Rogers, ESGR
Erioptera (Erioptera) chlorophylloides Alexander, 1919a: 106 - Rogers, ESGR
Erioptera (Erioptera) furcifera Alexander, 1919a: 108
Erioptera (Erioptera) gaspeana Alexander, 1929d: 250
Erioptera (Erioptera) georgei Alexander, 1956: 183 - Alexander, 1956
Erioptera (Erioptera) leptostyla Alexander, 1940: 154 - Alexander, 1940
Erioptera (Erioptera) megaphthalma Alexander, 1918: 60
Erioptera (Erioptera) septemtrionis Osten Sacken, 1859: 226 - Rogers, ESGR
Erioptera (Erioptera) straminea Osten Sacken, 1869: 157 - Rogers, ESGR
Erioptera (Erioptera) subfurcifera Alexander, 1929e: 51 - Alexander, 1929
Erioptera (Erioptera) uliginosa Alexander, 1930a: 277 - Rogers, ESGR
Erioptera (Erioptera) vespertina Osten Sacken, 1859: 226 - Rogers, ESGR

(syn.) Erioptera holoptica Dietz, 1921a: 245
(syn.) Erioptera fuscoantennata Dietz, 1921a: 246
Erioptera (Erioptera) villosa Osten Sacken, 1859: 226 - Rogers, ESGR
Erioptera (Mesocyphona) caliptera Say, 1823: 17 - Rogers, ESGR
Erioptera (Mesocyphona) knabi Alexander, 1913: 515
Erioptera (Mesocyphona) needhami Alexander, 1918: 383 - Rogers, ESGR
Erioptera (Mesocyphona) parva Osten Sacken, 1859: 227 - Rogers, ESGR
Erioptera (Hoplolabis) armata Osten Sacken, 1859: 227 - Rogers, ESGR
Erioptera (Psiloconopa) armillaris Osten Sacken, 1869: 158 - Rogers, ESGR
Erioptera (Psiloconopa) graphica Osten Sacken, 1859: 227 Rogers, ESGR
Erioptera (Psiloconopa) indianensis Alexander, 1922b: 1
Erioptera (Psiloconopa) venusta Osten Sacken, 1859: 227 - Alexander, 1956: 184 as ssp. nubilosa


Gonomyia Meigen, 1818: 146 (Tribe Eriopterini)
[saturated silt or sand at stream or pond margins]
Gonomyia (Idiocera) apicipina Alexander, 1926c: 117 - IN
Gonomyia (Idiocera) blanda Osten Sacken, 1859: 231 - Rogers, ESGR
Gonomyia (Idiocera) mathesoni Alexander, 1915b: 170
Gonomyia (Lipophleps) manca Osten Sacken, 1869: 178 - IN east
(syn.) Dicranomyia curvivena Coquillett, 1908: 144
Gonomyia (Lipophleps) sacandaga Alexander, 1914: 587
Gonomyia (Lipophleps) sulphurella Osten Sacken, 1859: 230 - Rogers, ESGR
Gonomyia (Neolipophleps) alexanderi (Johnson, 1912: 3 as Elliptera) - IL northeast
Gonomyia (Gonomyia) armigera Alexander, 1922b: 4 - IN northeast
Gonomyia (Gonomyia) bidentata Alexander, 1922b: 3 - IN, WI northeasst
Gonomyia (Gonomyia) cognatella Osten Sacken, 1859: 230 - WI east
Gonomyia (Gonomyia) florens Alexander, 1916c: 316 - Rogers, ESGR
Gonomyia (Gonomyia) kansensis Alexander, 1918: 158 - Rogers, ESGR
Gonomyia (Gonomyia) noveboracensis Alexander, 1916c: 319
Gonomyia (Gonomyia) reflexa Alexander, 1927d: 59 - Alexander, 1927d
Gonomyia (Gonomyia) subcinerea Osten Sacken, 1859: 231 - Rogers, ESGR

(syn.) Phyllolabis obscuris Doane, 1900: 192


Helius Lepeletier & Serville, in Latreille et al. 1828: 585 (Tribe Limoniini)
Helius flavipes (Macquart, 1855: 37 as Rhamphidia) - WI, widespread

(syn.) Limnobia prominens Walker, 1856: 435
(syn.) Rhamphidia brevirostris Osten Sacken, 1859: 222
Helius mainensis (Alexander, 1916a: 498 as Rhamphidia) - IL, WI, east


Hexatoma Latreille, 1809: 260 (Tribe Hexatomini)
[aquatic or semiaquatic]
Hexatoma (Hexatoma) magacera (Osten Sacken, 1859 as Anisomera) - IN, ON east
Hexatoma (Hexatoma) microcera Alexander, 1926b: 49 - NAm
Hexatoma (Eriocera) albitarsis (Osten Sacken, 1869: 357 as Penthoptera) - IW east
Hexatoma (Eriocera) cinerea (Alexander, 1912c: 169 as Eriocera) Rogers, ESGR
Hexatoma (Eriocera) fuliginosa (Osten Sacken, 1859: 243 as Eriocera)
Hexatoma (Eriocera) gibbosa (Doane, 1900: 193 as Eriocera) - Rogers, ESGR
Hexatoma (Eriocera) longicornis (Walker, 1848: 82 as Anisomera) - NAm
Hexatoma (Eriocera) spinosa (Osten Sacken, 1859: 244 as Arrhenica) - IL east
Hexatoma (Eriocera) tristis (Alexander, 1914: 602 as Eriocera) - IN east


Limonia Meigen, 1803: 262 (Tribe Limoniini)
Limonia (Limonia) globithorax (Osten Sacken, 1869: 74 as Dicranomyia) - WI-NF
Limonia (Limonia) indigena (Osten Sacken, 1859: 215 as Limnobia) - WI
Limonia (Limonia) macateei (Alexander, 1916b: 42 as Dicranomyia) - IN

(syn.) Dicranomyia varipes Dietz, 1921a: 241
Limonia (Limonia) maculicosta (Coquillett, 1905: 57 as Limnobia) - expect in northern UP
Limonia (Limonia) parietina (Osten Sacken, 1861: 289 as Limnobia)
Limonia (Limonia) sociabilis (Osten Sacken, 1869: 95 as Limnobia)
Limonia (Limonia) tristigma (Osten Sacken, 1859: 216 as Limnobia) - IL
Limonia (Dicranomyia) adirondacensis (Alexander, 1922a: 62 as Dicranomyia) - WI east
Limonia (Dicranomyia) brevivena (Osten Sacken, 1869: 66 as Dicranomyia) - widespread
Limonia (Dicranomyia) ctenopyga Alexander, 1943a: 156
Limonia (Dicranomyia) haeretica (Osten Sacken, 1869: 70 as Dicranomyia)

(syn.) Dicranomyia gibsoni Alexander, 1929b: 17
Limonia (Dicranomyia) halterata (Osten Sacken, 1869: 71 as Dicranomyia) - widespread

(syn.) Dicranomyia cinereipennis Lundström, 1912: 52
Limonia (Dicranomyia) halterella (Edwards, 1921: 201 as Dicranomyia)

(syn.) Dicranomyia gracilis Doane, 1900: 184
Limonia (Dicranomyia) humidicola (Osten Sacken, 1859: 210 as Dicranomyia)

(syn.) Dicranomyia viridicans Doane, 1908a: 7
(syn.) Dicranomyia badia, authors, not Walker
Limonia (Dicranomyia) immanis Alexander, 1950: 164
Limonia (Dicranomyia) immodesta (Osten Sacken, 1859: 211 - widespread
Limonia (Dicranomyia) immodestoides Alexander, 1919d: 327 - widespread, IN

(syn.) Dicranomyia iowensis Rogers, 1926: 50
Limonia (Dicranomyia) liberta (Osten Sacken, 1859: 209 as Dicranomyia) - WI, south and east
Limonia (Dicranomyia) longipennis (Schummel, 1829: 104 Limnobia) - widepsread
(syn.) Dicranomyia immomor Osten Sacken, 1861: 287
Limonia (Dicranomyia) michigana Alexander, 1950: 166
Limonia (Dicranomyia) moniliformis (Doane, 1900: 184 as Dicranomyia)
(syn.) Dicranomyia penicillata Alexander, 1927e: 7
(syn.) Dicranomyia haeretica authors, not Osten Sacken
Limonia (Dicranomyia) morioides (Osten Sacken, 1860: 17 as Dicranomyia)
Limonia (Dicranomyia) neomorio (Alexander, 1927d: 223 as Dicranomyia) - north widespread
Limonia (Dicranomyia) nycteris (Alexander, 1927d: 220 as Dicranomyia) - north widespread
Limonia (Dicranomyia) piscataquis Alexander, 1941: 86 - north widespread
(syn.) Dicranomyia vibei Nielsen, 1951: 185
Limonia (Dicranomyia) pudica (Osten Sacken, 1859: 212 as Dicranomyia)
Limonia (Dicranomyia) pudicoides Alexander, 1929a: 299 - IN
Limonia (Dicranomyia) rostifera (Osten Sacken, 1869: 65 as Dicranomyia) - north widespread
Limonia (Dicranomyia) sphagnicola (Alexander, 1925a: 173 as Dicranomyia) - widespread north and central
Limonia (Dicranomyia) spinifera (Alexander, 1927a: 229 as Dicranomyia)
Limonia (Dicranomyia) stulta (Osten Sacken, 1859: 210 as Dicranomyia) - WI east
(syn.) Furcomyia monticola Alexander, 1911b: 201
Limonia (Dicranomyia) terraenovae (Alexander, 1920c: 85 as Dicranomyia) - maybe north
(syn.) Limnobia teuipes Zetterstedt, 1838: 837
(syn.) Limnobia decora Staeger, 1840: 47
Limonia (Dicranomyia) uliginosa Alexander, 1929a: 27
Limonia (Geranomyia) canadensis (Westwood, 1835: 684 as Limnobiorhynchus) - widespread
Limonia (Geranomyia) communis (Osten Sacken, 1859: 207 as Geranomyia) - WI
Limonia (Geranomyia) distincta (Doane, 1900: 186 as Geranomyia)
Limonia (Geranomyia) diversa (Osten Sacken, 1859: 207 as Geranomyia) - widespread
Limonia (Geranomyia) rostrata (Say, 1823: 22 as Limnobia)
Limonia (unplaced subgenus) defuncta (Osten Sacken, 1859: 213 as Limnobia) - IL, WI widespread
Limonia (unplaced subgenus) fusca Meigen, 1804: 54
(syn.) Limnobia turpis Walker, 1856: 434
(syn.) Dicranomyia pubipennis Osten Sacken, 1859: 211
Limonia (unplaced subgenus) rara (Osten Sacken, 1869: 75 as Dicranomyia) - WI
Limonia (unplaced subgenus) shelfordi Alexander, 1944: 243


Limnophila Macquart, 1834: 95 (Tribe Hexatomini)
[wet soil ranging from sandy organic silt to black mud (Alexander in Stone et al. 1985)]
Limnophila (Lasiomastix) macrocera (Say, 1823: 20 as Limnobia) - Rogers, ESGR
Limnophila (Lasiomastix) subtenuicornis (Alexander, 1918: 61 as Lasiomastix) - Rogers, ESGR
Limnophila (Eutonia) marchandi Alexander, 1916d: 118
Limnophila (Prionolabis) munda Osten Sacken, 1869: 226 - WI east
Limnophila (Prionolabis) rufibasis Osten Sacken, 1859: 239 - Rogers, ESGR
Limnophila (Prionolabis) walleyi Alexander, 1929b: 187 - IL, ON east
Limnophila (Eloeophila) irene Alexander, 1927b: 58 - Alexander, 1927b
Limnophila (Eloeophila) solstitialis Alexander, 1926a: 109 - Rogers, ESGR
Limnophila (Dicranophragma) angustula Alexander, 1929b: 190 - Rogers, ESGR
Limnophila (Dicranophragma) fuscovaria Osten Sacken, 1859: 240 - Rogers, ESGR
Limnophila (Idioptera) fasciolata Osten Sacken, 1869: 206 - Rogers, ESGR
Limnophila (Phylidorea) adusta Osten Sacken, 1859: 235 - Rogers, ESGR
Limnophila (Phylidorea) adustoides Alexander, 1927b: 63 - IN east
Limnophila (Phylidorea) auripennis Alexander, 1926a: 113 - Rogers, ESGR
Limnophila (Phylidorea) caudifera Alexander, 1927b: 111 - Rogers, ESGR
Limnophila (Phylidorea) claggi Alexander, 1930b: 281
Limnophila (Phylidorea) consimilis Dietz, 1921a: 255 - Rogers, ESGR
Limnophila (Phylidorea) iowensis Alexander, 1927b: 112 - Rogers, ESGR
Limnophila (Phylidorea) luteola Alexander, 1927b: 113 - Rogers, ESGR
Limnophila (Phylidorea) persimilis Alexander, 1927b: 62 - IN
Limnophila (Phylidorea) platyphallus Alexander, 1926a: 111 - Rogers, ESGR
Limnophila (Phylidorea) siouana Alexander, 1929b: 188 - Rogers, ESGR
Limnophila (Phylidorea) subcostata (Alexander, 1911a: 287 as Phylidorea) - Rogers, ESGR
Limnophila (unplaced subgenus) brevifurca Osten Sacken, 1859: 237 - Rogers, ESGR
Limnophila (unplaced subgenus) laricicola Alexander, 1912c: 167 - Rogers, ESGR
Limnophila (unplaced subgenus) poetica Osten Sacken, 1869: 207 - Rogers, ESGR


Molophilus Curtis, 1833: pl. 444 (Tribe Eriopterini)
Molophilus cramptoni Alexander, 1924a: 61
Molophilus forcipulus (Osten Sacken, 1869: 163 as Erioptera) - Rogers, ESGR
Molophilus fultonensis Alexander, 1916a: 505 - Rogers, ESGR
Molophilus hirtipennis (Osten Sacken, 1859: 228 as Erioptera) - Rogers, ESGR
Molophilus huron Alexander, 1929e: 56 - Alexander, 1929; Rogers, ESGR
Molophilus novacaesariensis Alexander, 1916a: 506 - IN east
Molophilus pubipennis (Osten Sacken, 1859: 228 as Erioptera)
Molophilus quadrispinosus Alexander, 1924a: 62
Molophilus soror Alexander, 1927a: 230


Nasiternella Wahlgren, 1904: 4 (Tribe Pediciini)
Nasiternella hyperborea (Osten Sacken, 1861: 292 as Amalopsis) - widespread boreal


Ormosia Rondani, 1856: 180 (Tribe Eriopterini)
Ormosia (Scleroprocta) innocens (Osten Sacken, 1869: 142 as Rhypholophus)
Ormosia (Ormosia) arcuata (Doane, 1908b: 201 as Rhypholophus)
Ormosia (Ormosia) bilineata Dietz, 1916: 142 - Alexander, 1929 as O. huronis
(syn.) Ormosia huronis Alexander, 1929c: 20
Ormosia (Ormosia) broweri Alexander, 1939: 100 - widespread north
Ormosia (Ormosia) frisoni Alexander, 1920b: 224
Ormosia (Ormosia) hubbelli Alexander, 1926c: 20 - MN
Ormosia (Ormosia) ingloria Alexander, 1929c: 21 - IN, ON
Ormosia (Ormosia) luteola Dietz, 1916: 138
Ormosia (Ormosia) manicata (Doane, 1900: 187 as Rhypholophus) - Rogers, ESGR

(syn.) Ormosia deviata Dietz, 1916: 142
(syn.) Ormosia fuscopyga Alexander, 1924b: 3
Ormosia (Ormosia) meigenii (Osten Sacken, 1859: 226 as Erioptera)
Ormosia (Ormosia) monticola (Osten Sacken, 1869: 145 as Rhypholophus)
Ormosia (Ormosia) nigripila (Osten Sacken, 1869: 142 as Rhypholophus)
Ormosia (Ormosia) nimbipennis Alexander, 1917: 24
Ormosia (Ormosia) pygmaea (Alexander, 1912c: 166 as Trimicra)
Ormosia (Ormosia) romanovichiana Alexander, 1953: 327 - Rogers, ESGR
(syn.) Ormosia nubila Osten Sacken, 1859: 227 [nec. by Alexander, 1953: 327]
Ormosia (Ormosia) rubella (Osten Sacken, 1869: 144 as Rhypholophus) - Rogers, ESGR

(syn.) Ormosia enigmatica Dietz, 1921a: 249


Paradelphomyia Alexander, 1936: 184 (Tribe Hexatomini)
[organic silt along stream margins or in cool seepage areas (Alexander in Stone et al., 1965)]
Paradelphomyia (Oxyrhiza) americana (Alexander, 1912b: 829 as Adelphomyia)
Paradelphomyia (Oxyrhiza) cayuga (Alexander, 1912b: 831 as Adelphomyia)
(syn.) Ormosia abnormis Dietz, 1916: 137
(syn.) Adelphomyia hazletonensis Dietz, 1921a: 252
Paradelphomyia (Oxyrhiza) minuta (Alexander, 1912a: 286 as Adelphomyia)
Paradelphomyia (Oxyrhiza) pleuralis (Dietz, 1921a: 251 as Adelphomyia)


Phyllolabis Osten Sacken, 1877: 202 (Tribe Hexatomini)
Phyllolabis lagganensis Alexander, 1931: 183 - northern widespread


Pilaria Sintenis, 1889: 398 (Tribe Hexatomini)
[wet soil ranging from sandy organic silt to black mud (Alexander in Stone et al., 1965)]
Pilaria imbecilla (Osten Sacken, 1859: 237 as Limnophila) - ESGR, Rogers 1942
Pilaria meridiana (Staeger, 1840: 41 as Limnobia) - ESGR, Rogers 1942
(syn.) Limnophila osborni Alexander, 1914: 596
Pilaria quadrata (Osten Sacken, 1859: 241 as Limnophila) - ESGR, Rogers 1942
Pilaria recondita (Osten Sacken, 1869: 212 as Limnophila) - ESGR, Rogers 1942
Pilaria tenuipes (Say, 1823: 21 as Limnobia) - ESGR, Rogers 1942
(syn.) Limnophila edwardi Alexander, 1916a: 533

Prolimnophila Alexander, 1929b: 187 (Tribe Hexatomini)
[moist-to-saturated soil along streams or in marshes (Alexander in Stone et al. 1985)]
Prolimnophila areolata (Osten Sacken, 1859: 237 as Limnophila) - WI east


Pseudolimnophila Alexander, 1919c: 917 (Tribe Hexatomini)
[moist-to-saturated soil along streams or in marshes (Alexander in Stone et al. 1985)]
Pseudolimnophila australina Alexander, 1927d: 56 - IN east
Pseudolimnophila contempta (Osten Sacken, 1869: 218 as Limnophila)
Pseudolimnophila inornata (Osten Sacken, 1869: 219 as Limnophila)
Pseudolimnophila luteipennis (Osten Sacken, 1859: 236 as Limnophila)
Pseudolimnophila noveboracensis (Alexander, 1911b: 196 as Limnophila)


Tasiocera Skuse, 1890: 815 (Tribe Eriopterini)
Tasiocera (Dasymolophilus) niphadias (Alexander, 1925b: 229 as Molophilus)


Toxorhina Loew, 1850: 36 (Tribe Eriopterini)
[wet-to-saturated soil (Alexander in Stone et al. 1985)]
Toxorhina magna Osten Sacken, 1865: 232
Toxorhina muliebris Osten Sacken, 1865: 233 - Rogers, ESGR, not aquatic


Ulomorpha Osten Sacken, 1869: 232 (Tribe Hexatomini)
[wet soil ranging from sandy organic silt to black mud (Alexander in Stone et al. 1985)]
Ulomorpha pilosella (Osten Sacken, 1859: 241 as Limnophila)

PEDICIIDAE

Pedicia Latreille, 1809: 255 (Tribe Pediciini)
Pedicia (Pedicia) albivitta Walker, 1848: 37- MB-NF, MO-SC

(syn.) Pedicia goniphora Harris, 1835: 595
Pedicia (Pedicia) contermina Walker, 1848: 38
Pedicia (Pedicia) falcifera Alexander, 1941: 209
Pedicia (Tricyphona) autumnalis (Alexander, 1917: 30 as Tricyphona) - ON, WI east
Pedicia (Tricyphona) calcar (Osten Sacken, 1859: 247 as Amalopis) - WI east
Pedicia (Tricyphona) inconstans (Osten Sacken, 1859: 247 as Amalopis) - MN east
Pedicia (Tricyphona) katahdin (Alexander, 1914: 598 as Tricyphona)
Pedicia (Tricyphona) paludicola (Alexander, 1916a: 538 as Tricyphona)
Pedicia (Tricyphona) pumila Alexander, 1943b: 353

TIPULIIDAE

Tipula Linnaeus, 1758: 585 (Subfamily Tupuliinae, Tribe Tipulini)
Tipula (Nippotipula) abdominalis (Say, 1823: 18 as Ctenophora) - shredder, ubiquitous in streams and rivers in MI

(syn.) Tipula albilata Walker, 1848: 65
Tipula (Trichotipula) algonquin Alexander, 1915a: 469
Tipula (Oreomyza) angulata Loew, 1864: 61- Rogers, floodplains

(syn.) Tipula decora Doane, 1901: 125
Tipula (Schummelia) annulicornis Say, 1829: 151

(syn.) Tipula jejuna Johnson, 1909: 132
Tipula (Lunatipula) apicalis Loew, 1863: 277 (terrestrial)
Tipula (Vestiplex) balioptera Loew, 1863: 284 - WI
Tipula (Lunatipula) bicornis Forbes, 1890: 78 - Rogers, not aquatic but near marshes or marshes
Tipula (Oreomyza) borealis Walker, 1848: 66 - Rogers, floodplains

(syn.) Tipula hebes Loew, 1863: 285
Tipula (Yamatotipula) caloptera Loew, 1863: 292 - Rogers, aquatic in creek and brook gravel or sand
Tipula (Yamatotipula) cayuga Alexander, 1915a: 485 - Rogers, not aquatic
Tipula (Nobilotipula) collaris Say, 1823: 25
Tipula (Yamatotipula) concava Alexander, 1926b: 294
Tipula (Platytipula) cunctuans Say, 1823: 23 - Rogers, marsh edges (semiaquatic?)
Tipula (Yamatotipula) dejecta Walker, 1856: 442 - Rogers, semiaquatic marsh and seep edges

(syn.) Tipula fumosa Doane, 1901: 99
(nom. nud.) Tipula brevicollis Loew, 1863 in Alexander, 1915a: 460
Tipula (Yamatotipula) eluta Loew, 1863: 290 - Rogers, semiaquatic along marshes, creeks and seeps
Tipula (Arctotipula) dickinsoni Alexander, 1932: 240 - Rogers, semiaquatic marsh and seep edges
Tipula (Lunatipula) disjuncta Walker, 1856: 442 - Rogers, not aquatic

(syn.) Tipula taughannock Alexander, 1915a: 476

Tipula (Lunatipula) dorsimacula Walker, 1848: 69 - Rogers, not aquatic

(syn.) Tipula angustipennis Loew, 1863: 286
(syn.) Tipula beaulieui Dietz, 1921b: 301
(nom. nud.) Tipula nubilis Harris, 1835: 595
Tipula (Lunatipula) duplex Walker, 1848: 66 - Rogers, near marsh-wood margins, but not aquatic
Tipula (Pterelachisus) entomophthorae Alexander, 1918: 385 - not aquatic

(syn.) Tipula similissima Dietz, 1921a: 263
Tipula (Triplicitipula) flavoumbrosa Alexander, 1918: 415 - Rogers, not aquatic
Tipula (Yamatotipula) footeana Alexander, 1961: 11 - ID, NF
Tipula (Oreomyza) fragilis Loew, 1863: 279 - IN north and northeast
Tipula (Yamatotipula) fraterna Loew, 1864: 56 - Rogers, in algal mats of creeks
Tipula (Lunatipula) fuliginosa (Say, 1823: 18 as Ctenophora) - Rogers, not aquatic

(syn.) Tipula speciosa Loew, 1863: 288
Tipula (Vestiplex) fultonensis Alexander, 1918: 67 - northern
Tipula (Yamatotipula) furca Walker, 1848: 70 - Rogers, margins of creeks, marshes, semiaquatic in detritus

(syn.) Tipula bella Loew, 1863: 291
(syn.) Tipula fuscifer Harris in Johnson, 1925: 61
Tipula (Lunatipula) georgiana Alexander, 1915c: 133 - Rogers, not aquatic
Tipula (Oreomyza) helderbergensis Alexander, 1918: 64
Tipula (Schummelia) hermannia Alexander, 1915a: 480 - Rogers, semi-terrestrial (saturated soils)

(syn.) Tipula fasciata Loew, 1863: 279 [preocc. Linnaeus, 1767]
Tipula (Triplicitipula) hirsuta Doane, 1901: 113 - Doane, 1901; Rogers, not aquatic
Tipula (Oreomyza) huron Alexander, 1918: 66 - WI
Tipula (Oreomyza) illinoiensis Alexander, 1915c: 128 - Rogers, not aquatic
(syn.) Tipula versicolor Loew, 1863: 285 (preocc. Gmelin, 1792)
Tipula (Angarotipula) illustris Doane, 1901: 97 - Gelhaus 1986:136 (ESGR, Livingston Co., leg. S. Rogers, as Prionocera fuscipennis)

(syn.) Tipula fuscipennis Loew, 1866: 129
Tipula (Yamatotipula) kennicotti Alexander, 1915a: 480 - Rogers, marshes

(syn.) Tipula diluta Doane, 1901: 117 (preocc. Müller, 1764)
(syn.) Tipula parvemarginata Alexander, 1926b: 295
(syn.) Tipula tera Loew in Alexander, 1915a: 482
Tipula (Oreomyza) latipennis Loew, 1864: 60 - Rogers, floodplains, marsh margins

(syn.) Tipula ottawaensis Dietz, 1914: 349
Tipula (Vestiplex) longiventris Loew, 1863: 278 - Rogers, not aquatic
Tipula (Lunatipula) macrolabis - Loew, 1864: 58

(syn.) Tipula spectabilis Doane, 1901: 120
Tipula (Lunatipula) mainensis Alexander, 1915a: 475- ME-AB
(nom. nud.) Tipula laevigata Loew, in Alexander, 1915a: 476
Tipula (Lunatipula) megaura Doane, 1901: 112 - Doane, 1901; Rogers, wet margins of spring ponds and seeps, grassy streams
Tipula (Lunatipula) monticola Alexander, 1915a: 492 - Rogers, not aquatic
Tipula (Lunatipula) morrisoni Alexander, 1915a: 507 - IN east
Tipula (Nobilotipula) nobilis (Loew, 1864: 62 as Pachyrhina)
Tipula (Yamatotipula) noveboracensis Alexander, 1919b: 167
Tipula (Trichotipula) oropezoides Johnson, 1909: 131
Tipula (Trichotipula) pachyrhinoides Alexander, 1915a: 471 - MN
Tipula (Lunatipula) parshleyi Alexander, 1915a: 410 - Rogers, not aquatic
Tipula (Pterelachisus) penobscot Alexander, 1915a: 472 - not aquatic
Tipula (Oreomyza) phoroctenia Alexander, 1919b: 170 - ME west (expect in UP)
Tipula (unplaced) puncticornis Macquart, 1850: 319 - NAm
Tipula (Oreomyza) resurgens Walker, 1848: 67 - Rogers, floodplain

(syn.) Tipula grataLoew, 1863: 67
Tipula (Yamatotipula) sayi Alexander, 1911b: 194 - Rogers, inundated soils of marsh borders

(syn.) Tipula costalis Say, 1823
Tipula (Oreomyza) senega Alexander, 1915c: 128 - expect in UP
Tipula (Oreomyza) serta Loew, 1863: 283 - Doane, 1901 (as Tipula albonotata); Rogers, detritus of marsh and seep borders
(syn.) Tipula discolor Loew, 1863: 282
(syn.) Tipula albonotata Doane, 1901: 120
(syn.) Tipula ignota Alexander, 1915c: 128
Tipula (Trichotipula) stonei Alexander (for unifasciata (Loew, 1864: 64, as Pachyrhina) - Rogers, not aquatic

(syn.) Pachyrhina unifasciata Loew, 1863: 294
Tipula (Yamatotipula) strepens Loew, 1863: 291 - Rogers, semiaquatic, moss and algal mats of rills and seeps
Tipula (Oreomyza) subfasciata Loew, 1863: 282 - WI
Tipula (Lunatipula) submaculata Loew, 1863: 288 - Rogers, not aquatic

(syn.) Tipula cuspidata Doane, 1901: 111
(nom. nud.) Tipula bidens Loew in Alexander, 1915a: 464
Tipula (Yamatotipula) sulphurea Doane, 1901: 99 - Rogers, semiaquatic, shallow semiflowing pools between marsh hummocks
Tipula (Yamatotipula) tephrocephala Loew, 1864: 62 - Rogers, seepages, wet soil, but not aquatic
Tipula (Lunatipula) translucida Doane, 1901: 109 - Rogers, floodplains, not aquatic
Tipula (Yamatotipula) tricolor Fabricius, 1775: 749 - Rogers, marshes, streams, seepages and stagnant pools

(syn.) Tipula vitrea Wulp, 1881: 150
Tipula (Triplicitipula) triplex Walker, 1848: 66 - Doane, 1901 (as Tipula inermis); Rogers, open floodplains, not aquatic

(syn.) Tipula inermis Doane, 1901: 112
Tipula (Oreomyza) trivitata Say, 1823: 26 - Rogers, floodplains, swamp margins
Tipula (Trichotipula) unimaculata (Loew, 1864: 64 as Pachyrhina) - Rogers, floodplain woods, seepages, not aquatic
Tipula (Platytipula) ultima Alexander, 1915c: 128 - Rogers, margins of marshes

(syn.) Tipula flavescens Fabricius, 1805: 24
(syn.) Tipula flavicans Fabricius, 1805: 373
Tipula (Triplicitipula) valida Loew, 1863: 287 - Doane, 1901 (as Tipula calva); Rogers, floodplain woods, not aquatic

(syn.) Tipula calva Doane, 1901: 114
Tipula (Yamatotipula) vicina Dietz, 1917: 148 - Rogers, Stone - sand and gravel bottom banks
Tipula (Lunatipula) youngi Alexander, 1927c: 218 - Rogers, seepage areas, flood-plains

unplaced Tipulidae species
Limnobia ignobilis Walker, 1856: 435, nomen dubium, holotype lost

References - Distribution

Kovalak WP. 1978. Diel changes in stream benthos density on stones and artificial substrates. Hydrobiologia 58(1): 7-16.
Rogers JS. 1942. The Crane Flies (Tipulidae) of the George Reserve, Michigan. Miscellaneous Publications of the Museum of Zoology, University of Michigan 53: 129 pp., 8 pl. PDF.

References - Taxonomy

Alexander CP. 1911a. New Tipulidae (Diptera). The Canadian Entomologist 43: 286-288. PDF.
Alexander CP. 1911b. Synonymical, and other notes on the Tipulidae (Diptera). Psyche 18: 192-203. PDF.
Alexander CP. 1912a. On the tropical American Rhipidiae (Tipulidae, Dipt.). Bulletin of the Brooklyn Entomological Society 8: 6-17. PDF.
Alexander CP. 1912b. The American species of Adelphomyia Bergroth (Tipulidae, Dipt.). Pomona College Journal of Entomology 4: 829-831. PDF.
Alexander CP. 1912c. New Nearctic Tipulidae (Diptera). Psyche 19: 163-171. PDF.
Alexander CP. 1913. A synopsis of part of the Neotropical crane-flies of the subfamily Limnobinae. Proceedings of the United States National Museum 44: 481-549. PDF.
Alexander CP. 1914. New or little-known craneflies from the United States and Canada. Tipulidae, Diptera. Proceedings of the Academy of Natural Sciences of Philadelphia 66: 579-606. PDF.
Alexander CP. 1915a. New or little-known crane-flies from the United States and Canada: Tipulidae, Diptera. Part. 2. Proceedings of the Academy of Natural Sciences of Philadelphia 67: 458-514. PDF.
Alexander CP. 1915b. A new Nearctic Gonomyia (Tipulidae, Diptera). Entomological News 26: 170-172. PDF.
Alexander CP. 1915c. New Nearctic crane-flies in the United States national museum (Diptera, Tipulidae). Insecutor Inscitiae Menstruus 3: 127-142. PDF.
Alexander CP. 1916a. New or little-known crane-flies from Colombia, Ecuador and Peru (Tipulidae, Diptera). Transactions of the American Entomological Society 42: 1-32. PDF.
Alexander CP. 1916b. New Nearctic crane-flies (Tipulidae, Diptera). The Canadian Entomologist 48: 42-53. PDF.
Alexander CP. 1916c. New North American species of the genus Gonomyia Meigen (Tipulidae, Diptera). The Canadian Entomologist 48: 316-325. PDF.
Alexander CP. 1916d. New limnophiline crane-flies from the United States and Canada (Tipulidae, Diptera). Journal of the New York Entomological Society 24: 118-125. PDF.
Alexander CP. 1917. New Nearctic crane-flies (Tipulidae, Diptera). Part II; Part III. The Canadian Entomologist 49: 22-31 (PDF), 61-64 (PDF); 199-211 p. 12. (PDF).
Alexander CP. 1918. New Nearctic crane-flies (Tipulidae, Diptera). Part IV; Part V; Part VI. The Canadian Entomologist 50: 60-71 (PDF); 158-165 (PDF), 242-246, 1 fig. (PDF); 381-386 (PDF), 411-416 (PDF).
Alexander CP. 1919a. New Nearctic species of the genus Erioptera Meigen (Tipulidae, Diptera). Bulletin of the Brooklyn Entomological Society 14: 104-108. PDF.
Alexander CP. 1919b. New Nearctic crane-flies (Rhyphidae and Tipulidae, Diptera). Part VII. The Canadian Entomologist 51: 162-172. PDF.
Alexander CP. 1919c. The crane-flies of New York. Part I. Distribution and taxonomy of the adult flies. Memoirs, Cornell University Agricultural Experiment Station 25: 767-993. PDF.
Alexander CP. 1920a. Two undescribed pediciine crane-flies from the United States (Tipulidae, Diptera). The Canadian Entomologist 52: 78-80. PDF.
Alexander CP. 1919d. Undescribed species of Japanese crane-flies (Tipulidae, Diptera). Annals of the Entomological Society of America 12: 327-348. PDF.
Alexander CP. 1920b. New Nearctic crane-flies (Tipulidae, Diptera). Part IX. The Canadian Entomologist 52: 109-112. PDF.
Alexander CP. 1920c. New species of crane-flies from the United States and Canada (Tipulidae, Diptera). Journal of Entomology and Zoology 12: 85-92.
Alexander CP. 1922a. The crane-flies of New York: first supplementary list. Bulletin of the Brooklyn Entomological Society 17: 58-62. PDF.
Alexander CP. 1922b. New species of crane-flies from southern Indiana. Occasional Papers of the Museum of Zoology, University of Michigan 127: 1-7. PDF.
Alexander CP. 1924a. The crane-flies of New York: second supplementary list. Bulletin of the Brooklyn Entomological Society 19: 57-64. PDF.
Alexander CP. 1924b. New species of two-winged flies from western North America belonging to the family Tipulidae. Proceedings of the United States National Museum 64(10): 1-16. PDF.
Alexander CP. 1925a. The crane-flies (Tipulidae) of New England: first supplementary list. Occasional Papers of the Boston Society of Natural History 5: 169-174. PDF.
Alexander CP. 1925b. Undescribed species of crane-flies from the eastern United States and Canada (Dipt.: Tipulidae). Part I [concl.]. Entomological News 36: 229-230. PDF.
Alexander CP. 1926a. Undescribed species of the genus Limnophila from eastern North America (Tipulidae, Diptera). Part I. Bulletin of the Brooklyn Entomological Society 21: 109-115. PDF.
Alexander CP. 1926b. Undescribed species of crane-flies from the eastern United States and Canada (Dipt.: Tipulidae). Part II. Entomological News 37: 44-51. PDF.
Alexander CP. 1926c. Undescribed species of crane-flies from the United States and Canada. Part II. (Diptera, Tipulidae). Insecutor Inscitiae Menstruus 14: 114-122. PDF.
Alexander CP. 1927a. The crane-flies (Tipulidae) of New England: second supplementary list. Occasional Papers of the Boston Society of Natural History 5: 223-231. PDF.
Alexander CP. 1927b. Undescribed species of the genus Limnophila from eastern North America (Tipulidae, Diptera). Part II. Bulletin of the Brooklyn Entomological Society 22: 56-64. PDF.
Alexander CP. 1927c. Records and descriptions of crane-flies from Alberta (Tipulidae, Diptera). I. The Canadian Entomologist 59: 214-225. PDF.
Alexander CP. 1927d. Records and descriptions of crane-flies from the eastern United States (Tipulidae, Diptera). Journal of the New York Entomological Society 35: 55-63. PDF.
Alexander CP. 1927e. Undescribed crane-flies from the Holarctic region in the United States national museum. Proceedings of the United States National Museum 72(2): 1-17. PDF.
Alexander CP. 1929a. The crane-flies of New York: third supplementary list. Bulletin of the Brooklyn Entomological Society 24: 22-29. PDF.
Alexander CP. 1929b. Undescribed species of the genus Limnophila from eastern North America (Tipulidae, Diptera). Part IV. Bulletin of the Brooklyn Entomological Society 24: 187-191. PDF.
Alexander CP. 1929c. New Nearctic crane-flies (Tipulidae, Diptera). Part XIII. The Canadian Entomologist 61: 15-22. PDF.
Alexander CP. 1929d. A list of the crane-flies of Quebec (Diptera). I [part]. The Canadian Entomologist 61: 231-236. PDF.
Alexander CP. 1929e. Undescribed species of eriopterine crane-flies from the United States and Canada (Tipulidae, Diptera). Part I. Journal of the New York Entomological Society 37: 49-58. PDF.
Alexander CP. 1930a. The crane-flies (Tipulidae) of New England: third supplementary list. Occasional Papers of the Boston Society of Natural History 5: 267-278. PDF.
Alexander CP. 1930b. New or insufficiently-known crane-flies from the Nearctic region (Tipulidae, Diptera). Part I; Part II. Bulletin of the Brooklyn Entomological Society 25: 71-77 (PDF), 276-282.
Alexander CP. 1931. New or insufficiently-known crane-flies from the Nearctic region (Tipulidae, Diptera). Part III. Bulletin of the Brooklyn Entomological Society 26: 177-184. PDF.
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Alexander CP. 1936. New or little-known Tipulidae from eastern Asia (Diptera). XXX. Philippine Journal of Science 60: 165-204. PDF.
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Alexander CP. 1940. New Nearctic crane-flies (Tipulidae, Diptera). Part XIV. The Canadian Entomologist 72: 151-155. PDF.
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April 10, 2017

• Corrections to nomenclatures and organization, splitting references into sections for distribution and taxonomy.

Page created: November 20, 2002 - Last updated: May 18, 2017 (EB)