Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Chironomidae Newman, 1834: 379 - Non-biting midges

Chironomidae are certainly the most speciose and frequently most numerous insect family in most freshwater systems, and are often integral energy-processing components of almost all freshwater ecosystems. The family also has a number of semi-terrestrial and terrestrial species.

The systematics and taxonomy of such a large group is still in a state of change, with relatively few expert taxonomists active in describing the enormous diversity of this group. Many groups in Chironomidae remain poorly understood, especially genetically, and undoubtedly there exist within genera species-complexes that are only distinguishable with various molecular methods (e.g., Martin 2003). Further, some taxa simply have been poorly sampled and studied, and undoubtedly many new species await description. A joint project by Pat Hudson and Ethan Bright to survey the Chironomidae of the Great Lakes Region, with an emphasis on Michigan, is a new start to systematically update our information. The following list must be considered preliminary and by no means definitive. Distribution records are often based on few observations, and for few groups are we knowledgeable about ecological parameters from which we can infer a distribution. Thus, one must be liberal in listing species that could occur in our state. Currently, there are 305 species that have been documented from Michigan, with at least another 247 species (not including undescribed forms) likely to occur here (E. Bright, 2017, unpub. info.).

Genera are denoted in green; species denoted in bold blue have been recorded in Michigan; other species denoted in black bold are expected to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

    Subfamily CHIRONOMINAE

      Acalcarella Shilova, 1955: 319 (Tribe Chironomini)

        Acalcarella sp. - unidentified larvae collected from from unspecified location in Lake Superior (Stroom et al. 2003)

      Apedilum Townes, 1945: 32 (Tribe Chironomini)

        Apedilum elachistus Townes, 1945: 33 - Oliver et al. 1990:40
        Apedilum subcinctum Townes, 1945: 33 - ONT-CA, NM-FL (Townes 1945:33; Oliver et al. 1990)

      Axarus Roback, 1980: 32 (Tribe Chironomini)

        Axarus festivus (Say, 1823: 13 as Chironomus) - UMMZ
          (syn.) Chironomus lineatus Say, 1823: 14
          (syn.) Chironomus lineola Say, 1823, emend. Wiedemann, 1828: 16-17
          (syn.) Chironomus lasiomerus Walker, 1848:19 (comb.) Xenochironomus festivus (Say), Townes 1945: 94-95
        Axarus scopula (Townes, 1945: 93-94 as Xenochironomus) - Townes 1945: 93-94
        Axarus taenionotus (Say, 1829: 149, as Chironomus) - Oliver et al. 1990: 41
          (comb.) Xenochironomus taenionotus (Say), Townes 1945: 95

      Beckidia Sæther, 1979: 315 (Tribe Chironomini)

        Beckidia tethys (Townes, 1945: 152 as Harnischia (Cladopelma)) - ON, widespread north, rockies

      Benthalia Lipina, 1939: 107 (Tribe Chironomini)

        [note: Jon Martin places these species in Benthalia; see http://www.genetics.unimelb.edu.au/Martin/NACytfiles/Sp3k.htm
        Benthalia brunneipennis (Johannsen, 1905: 205 as Chironomus) - Townes 1945:112 (as Tendipes (Einfeldia) brunneipennis); Jon Martin (accessed Feb. 2014)
        Benthalia natchitocheae (Sublette, 1964: 124 as Chironomus (Chironomus)) - Hudson and Bright 2012
          (comb.) Chironomus (Lobochironomus) natchitocheae Sublette, 1964: 124

      Chernovskiia Sæther, 1977: 107 (Tribe Chironomini)

        Chernovskiia amphitrite (Townes, 1945: 151 as Harnischia (Cladopelma)) - widespread
        Chernovskiia orbicus (Townes, 1945: 151-152 as Harnischia (Cladopelma)) - widespread

      Chironomus Meigen, 1803: 261 (Tribe Chironomini)

        Chironomus (Chironomus) aberratus Keyl, 1961: 484 - Egan & Ferrington 2015: 13 (Isle Royale, pupal exuviae)
        Chironomus (Chironomus) anthracinus Zettersted, 1860: 6499 - Townes 1945:130-131, Egan & Ferrington 2015: 14 (Isle Royale, pupal exuviae)
          (syn.) Chironomus rempelii Zetterstedt, 1860: 6499
          (syn.) Chironomus hyperboreus authors, not Staeger
        Chironomus (Chironomus) atrella (Townes, 1945: 130 as Tendipes (Tendipes)) - UMMZ
        Chironomus (Chironomus) atritibia Malloch, 1934: 16 - ON, widepsread north (Townes 1945:131, Saether 1970:7; Oliver et al. 1990:42
        Chironomus (Chaetolabis) atroviridis (Townes, 1945: 114 as Tendipes (Tendipes)) - Townes 1945:114-115; Jon Martin (accessed Feb. 2014)
        Chironomus (Chironomus) bifurcatus Wülker et al. 2009:36 - Wülker et al. 2009: 36-44; Saether 2009:5-8; Jon Martin (accessed Feb. 2014)
          (syn.) Chironomus species a Martin 1979
          (syn.) Chironomus decorus-group species 1 Butler et al. 1995
          (syn.) Chironomus species a Martin 2008
        Chironomus (Chironomus) crassicaudatus Malloch, 1915: 453 - (Midland Co.) Townes 1945: 134-135
          (comb.) Tendipes (Tendipes) crassicaudatus (Malloch), Townes 1945: 134
        Chironomus (Chironomus) cucini Webb, 1969: 105 - IN, ON (Oliver et al. 1990: 42), WI (Butler 1987)
        Chironomus (Chironomus) decorus Johannsen, 1905: 239 - Townes 1945: 120-123
          (syn.) Chironomus similis Johannsen, 1905: 236
          (syn.) Chironomus coyugae Johannsen in Tilbury, 1913: 308
          (syn.) Chironomus lasiopus Walker, 1848: 19 (nom. dubia)
          (syn.) Chironomus redeuns Walker, 1856b: 422 (nom. dubia)
          (syn.) Chironomus distinguendus Kieffer, 1917:346
          (nom.dub.) Chironomus attenuatus Walker, 1848:20
        Chironomus (Chironomus) dilutus Shobanov et al. 1999: 315 - Martin 2012 (http://www.genetics.unimelb.edu.au/martin/NACytfiles/Spt.html)
          (syn.) Chironomus pallidivittatus Malloch, 1915: 445 (not sensu Edwards 1929)
        Chironomus (Chironomus) entis Shobanov, 1989 - Saginaw Bay (Jon Martin (accessed Feb. 2014))
          Note: this is a sibling species of C. plumosus, cytologically determined
        Chironomus (unplaced) harti Malloch, 1915: 457 - IL
        Chironomus (Chironomus) hyperboreus Staeger, 1845: 349 - UMMZ
        Chironomus (Chironomus) maturus Johannsen, 1908: 279 - IN, widespread (Oliver et al. 1990: 42)
        Chironomus (Chironomus) melanescens Keyl, 1962: 472 - ON, WI (Jon Martin, pers.comm., April 2015)
          Chironomus (Chironomus) sp. e Martin, webpage
        Chironomus (Chironomus) mozleyi Wülker, 2007: 322
        Chironomus (Chaetolabis) ochreatus (Townes, 1945: 115 as Tendipes) - Townes 1945:115; Jon Martin (accessed Feb. 2014)
        Chironomus (Camptochironomus) pallidivittatus (Malloch, 1915: 455 as Chironomus tentans var. pallidivittatus)
        Chironomus (Chironomus) plumosus (Linneaus, 1758: 587 as Tipula) - Townes 1945; Jon Martin (accessed Feb. 2014)
          (syn.) Tipula annularis De Geer, 1776: 379
          (syn.) Chironomus cristatus Fabricius, 1805: 39
          (syn.) Chironomus ferrugineovittatus Zetterstedt, 1850: 3492
          (syn.) Chironomus imperator Walley, 1926b: 64
        Chironomus prasinus Meigen, 1818 - Kiknadze et al. 2016: 332
          (syn.) Chironomus vancouveri Michailova and Fischer, 1986: 99
        Chironomus (Chironomus) riparius Meigen, 1804: 13 - Needham 1908: 255-257; Townes 1945
          (syn.) Chironomus albistria Walker, 1848: 17
          (syn.) Chironomus serus Malloch, 1915: 481
          (syn.) Chironomus militaris Johannsen, 1937b: 46
          (syn.) Tendipes thummi Kieffer, 1911a: 36
        Chironomus (Chironomus) staegeri Lundbeck, 1898: 271 - Townes 1945; Jon Martin (accessed Feb. 2014)
        Chironomus (Chironomus) stigmaterus Say, 1823: 15 - widespread
          (syn.) Chironomus glaucurus Wiedemann, 1828: 15
        Chironomus (Chironomus) tuxis Curran, 1930: 31 (Tendipes (Tendips) (orig.) - IL, MN, ON, widespread (Townes, 1945: 123-124; however, Saether (2012) lists it in Michigan)
        Chironomus (Chironomus) winnelli Wülker, 2007: 325 - Lake Michigan (type locality, 43.001N, -86.225W est.) (Wülker, 2007)

      Cladopelma Kieffer, 1921b: 274 (Tribe Chironomini)

        Cladopelma amachaerum (Townes, 1945 as Harnischia) - new record (Saline River in Milan: 42.0824, -83.6841; Bright & Hudson, in prep.)
        Cladopelma collator (Townes, 1945 as Harnischia) - new record (Silver Lake near Pinckney: 42.4161,-83.9614; Bright & Hudson, in prep.)
        Cladopelma edwardsi (Kruseman, 1933: 194 as Tendipes) - Hudson & Bright 2012
          (comb.) Parachironomus edwardsi (Kruseman, 1933), Brundin 1947: 55
          (comb.) Harnischia (Harnischia) edwardsi (Kruseman, 1933), Townes 1945: 167
          (comb.) Chironomus (Cryptochironomus) edwardsi (Kruseman, 1933), Sublette 1964: 134
          (comb.) Cryptocladopelma edwardsi (Kruseman, 1933), Brundin 1949: 741
          (syn.) Chironomus (Chironomus) virescens Meigen, 1818 sensu Edwards (1929: 391)
          (syn.) Harnischia viridula auct. nec C. viridulum (Linnaeus) Wang et al. (1977: 230)
          (syn.) Cladopelma indicum Bhattacharyay, Duta & Chaudhuri, 1985: 179
          (syn.) Cryptotendipes fujiquartus Sasa, 1985: 109
          (syn.) Cladopelma fujiquarta Sasa, 1993: 71
          (syn.) Cryptotendipes sibaabeus Sasa, Sumita & Suzuki, 1999: 184
          (syn.) Cryptotendipes inawaabeus Sasa, Kitami & Suzuki, 1999: 8
          (syn.) Harnischia daitoheia Sasa & Suzuki, 2001a: 70
          (syn.) Harnischia ginzandeeus Sasa & Suzuki, 2001b: 12
          (syn.) Harnischia inadeeus Sasa, Kitami & Suzuki, 2001: 5
          (syn.) Harnischia sibacedea Sasa, Sumita & Suzuki, 1999: 185
          (syn.) Harnischia simantocedea Sasa, Suzuki & Sakai, 1998: 50
        Cladopelma galeator (Townes, 1945: 170 as Harnischia) - new records (Saline River in Milan 42.0824,-83.6841; Huron River below Ford Lake 42.2095,-83.5404; Bright & Hudson, in prep.)
        Cladopelma viridulum (Linnaeus, 1767: 975 as Tipula) - Cook-MSU
          (comb.) Chironomus (Harnischia) viridulus (Linnaeus) Goetghebuer 1928: 86
          (comb.) Chironomus (Chironomus) viridulus (Linnaeus) Edwards 1929: 390
          (comb.) Chironomus (Cryptochironomus) viridulus (Fabricius, 1805), Tokunaga 1940: 301
          (comb.) Harnischia (Harnischia) viridulus (Linnaeus) Townes 1945: 34
          (comb.) Cryptocladopelma viridula (Linnaeus) Reiss 1968: 199
          (comb.) Cladopelma viridula (Linnaeus) Yamamoto 1997: 585

      Cladotanytarsus Kieffer, 1921b: 277 (Tribe Tanytarsini)

        Cladotanytarsus aeiparthenus Bilyj, 1989: 948 - ONT
        Cladotanytarsus daviesi Bilyj, 1989: 951 - ONT
        Cladotanytarsus elaensis Bilyj, 1989: 955 - Hudson & Bright 2012
        Cladotanytarsus fusiformis Bilyj, 1989: 954 - ONT, OH
        Cladotanytarsus muricatus Bilyj, 1989: 956 - ONT
        Cladotanytarsus pinnaticornis Bilyj, 1989: 953 - ONT
        Cladotanytarsus tribelus Bilyj, 1989: 957 - ONT
        Cladotanytarsus viridiventris (Malloch, 1915: 491 as Tanytarsus) - Malloch, 1915

      Constempellina Brundin, 1947: 82 (Tribe Tanytarsini)

        Constempellina brevicosta (Edwards, 1937: 146 as Tanytarsus.) - widespread north

      Corynocera Zetterstedt, 1838 (Tribe Tanytarsini)

        Corynocera sp. - a Holarctic genus, records of undescribed species are reported from North Dakota (Malcolm Butler, pers.comm.) and Nebraska (Barbara Hayford, pers.comm.), indicating a much more widespread distribution, and may be encountered in our state

      Cryptochironomus Kieffer, 1918a: 46 (Tribe Chironomini)

        Cryptochironomus blarina Townes, 1945: 99 - AB-ONT, MN-NY, s. to FL, OH
        Cryptochironomus conus Mason, 1986: 400 - SK-ONT, OH
        Cryptochironomus curryi Mason 1986: 404 - Saether 2009; Curry 1958 (as C. fulvus)
          (misid.) Cryptochironomus fulvus (Johannsen) Curry 1958: 435
        Cryptochironomus digitatus (Malloch, 1915: 483 as Chironomus) - Townes 1945
        Cryptochironomus fulvus (Johannsen, 1905: 224 as Chironomus) - Townes 1945, UMMZ
          (syn.) Chironomus mallochi Kieffer, 1919b: 191 (n. nam. for Chironomus abbreviatus Malloch, 1915b: 451, preocc. Kieffer, 1913a)
          (syn.) Chironomus parvilamellatus Malloch, 1915: 479
        Cryptochironomus ponderosus (Sublette, 1964b: 129 as Chironomus) - SK-OH, s. to SC
        Cryptochironomus psittacinus (Meigen, 1830: 247 as Chironomus) - Townes 1945
          (syn.) Chironomus obtusilobus Malloch, 1923: 171
          (syn.) Chironomus farinalis Walley, 1926b: 205
        Cryptochironomus scimitarus Townes, 1945: 98 - Townes 1945
        Cryptochironomus sorex Townes, 1945: 97 - Oliver et al. 1990:44
        Cryptochironomus stylifera Johannsen, 1908: 281, Sæther 2009 - Townes 1945

      Cryptotendipes Lenz, 1941: 34 (Tribe Chironomini)

        Cryptotendipes casuarius (Townes, 1945: 162 as Harnischia) - ON, widespread
        Cryptotendipes darbyi (Sublette, 1960: 221 as Tendipes) - OH, widespread
        Cryptotendipes emorsus (Townes, 1945: 161 as Harnischia) - OH, widespread eastern NA
        Cryptotendipes pseudotener Goetghebuer, 1922: 38 - Townes 1945: 164, Oliver et al. 1990: 44

      Cyphomella Sæther, 1977: 101 (Tribe Chironomini)

        Cyphomella grisea (Malloch, 1915: 468 as Chironomus) - Oliver et al. 1990: 45

      Demeijerea Kruseman, 1933: 154 (Tribe Chironomini)

        Demeijerea atrimana (Coquillett, 1902b: 94 as Chironomus) - Cook-MSU
        Demeijerea brachialis (Coquillett, 1901b: 607 as Chironomus) - Cook-MSU
        Demeijerea obreptus (Townes, 1945: 139 as Glyptochironomus) - UMMZ

      Demicryptochironomus Lenz, 1941: 34 (Tribe Chironomini)

        Demicryptochironomus (Irmakia) cuneatus (Townes, 1945: 163 as Harnischia) - Cook-MSU
        Demicryptochironomus (Irmakia) fastigatus (Townes, 1945: 162 as Harnischia) - Oliver et al. 1990:45

      Dicrotendipes Kieffer, 1913d: 23 (Tribe Chironomini)

        Dicrotendipes botaurus (Townes, 1945: 109 as Tendipes) - Townes 1945: 109-110
        Dicrotendipes fumidus (Johannsen, 1905: 221 as Chironomus) - Townes 1945: 104, Egan & Ferrington 2015: 15 (Isle Royale, pupal exuviae)
          (syn.) Chironomus incognitus Malloch, 1915
        Dicrotendipes leucoscelis (Townes, 1945: 104 as Tendipes) - Townes 1945: 104-105
        Dicrotendipes lobiger (Kieffer, 1921a: 71 as Limnochironomus) - Townes 1945: 103-104
        Dicrotendipes lucifer (Johannsen, 1907: 110 as Chironomus) - OH, widespread
          (syn.) Chironomus indistinctus Malloch, 1915: 477
        Dicrotendipes milleri (Townes, 1945: 110 as Tendipes) - Townes 1945: 110
        Dicrotendipes modestus (Say, 1823: 13 as Chironomus) - Townes 1945: 106-107, Egan & Ferrington 2015: 15 (Isle Royale, pupal exuviae)
        Dicrotendipes neomodestus (Malloch, 1915: 475 as Chironomus) - Townes 1945: 107
          (syn.) Chironomus tenellus authors, not Zetterstedt
        Dicrotendipes nervosus (Staeger, 1839: 567 as Chironomus) - Townes 1945: 108-109, Egan & Ferrington 2015: 15 (Isle Royale, pupal exuviae)
          (syn.) Chironomus brevitibialis Zetterstedt, 1850
        Dicrotendipes tritomus (Kieffer, in Thienemann & Kieffer, 1916: 524 as Chironomus) - AK-NH, s. to LA-FL, OH
          (syn.) Chironomus incurvus Sublette, 1946b: 126

      Einfeldia Kieffer, 1924a: 393 (Tribe Chironomini)

        Einfeldia chelonia (Townes, 1945: 114 as Tendipes) - OH, widespread east USA
        Einfeldia pagana (Meigen, 1838: 7 as Chironomus) - Oliver et al. 1990: 46; Jon Martin (accessed Feb. 2014)
          (syn.) Einfeldia synchrona Oliver, 1971: 1591

      Endochironomus Kieffer, 1918b: 69 (Tribe Chironomini)

        Endochironomus nigricans (Johannsen, 1905: 219 as Chironomus) - Townes 1945: 64-65; Berg 1950: 97-98; Grodhaus 1987, Egan & Ferrington 2015: 15 (Isle Royale, pupal exuviae)
          (syn.) Chironomus johnsoni Kieffer, 1906a (n. nam. for Chironomus albipennis Meigen of Johnson, 1900, nom. nud.).
          (syn.) Chironomus albipennis authors, not Meigen
        Endochironomus subtendens (Townes, 1945: 65 as Tanytarsus) - Townes 1945

      Gillotia Kieffer, 1921b: 272 (Tribe Chironomini)

        Gillotia alboviridis (Malloch, 1915: 482 as Chironomus) - IL, OH (maybe extreme s. LP)

      Glyptotendipes Kieffer, 1913: 255 (Tribe Chironomini)

        Glyptotendipes amplus Townes, 1945: 145 - widespread north
        Glyptotendipes barbipes (Staeger, 1839: 561 as Chironomus) - Townes 1945
        Glyptotendipes dreisbachi Townes 1945: 145 - Townes 1945; Berg 1950: 94-96
        Glyptotendipes lobiferus (Say, 1823: 12 as Chironomus) - Townes 1945; Berg 1950: 92-94
          (syn.) Chironomus ithacanensis Johannsen, 1908: 279
          (syn.) Glyptotendipes americanus Kieffer, 1917: 355
        Glyptotendipes meridionalis Dendy & Sublette, 1959: 517 - OH, widespread
        Glyptotendipes paripes (Edwards, 1929: 392 as Chironomus) - Townes 1945
        Glyptotendipes senilis (Johannsen, 1937b: 37 as Chironomus) - Townes 1945
        Glyptotendipes unacus Townes 1945: 146 - widespread north

      Goeldichironomus Fittkau, 1965: 210 (Tribe Chironomini)

        Goeldichironomus holoprasinus (Goeldi, 1905: 135 as Chironomus) - OH, widespread
          (syn.) Chironomus fulvipilus Rempel, 1939: 210

      Harnischia Kieffer, 1921b: 273 (Tribe Chironomini)

        Harnischia cutilamellata (Malloch, 1915: 474 as Chironomus) - Malloch, 1915

      Hyporhygma Reiss, 1982: 289 (Tribe Chironomini)

        Hyporhygma quadripunctatum (Malloch, 1915: 437 as Chironomus) - Cook-MSU

      Kiefferulus Goetghebuer, 1922: 40 (Tribe Chironomini)

        Kiefferulus (Kiefferulus) dux (Johannsen, 1905: 231 as Chironomus) - Townes 1945, UMMZ

      Kloosia Kruseman, 1933: 152 (Tribe Chironomini)

        Kloosia sp. - unassociated larvae collected from unspecified location in Lake Superior (Stroom et al. 2003)

      Lauterborniella Thienemann & Bause, 1913: 120 (Tribe Chironomini)

        Lauterborniella agrayloides (Kieffer, 1911a: 51 as Tanytarsus) - Hudson & Bright 2012

      Lipiniella Shilova, 1961: 19 (Tribe Chironomini)

        Lipiniella sp. - unidentified larvae collected from unspecified location in Lake Superior (Stroom et al. 2003)

      Microchironomus Kieffer, 1918b: 113 (Tribe Chironomini)

        Microchironomus nigrovittatus (Malloch, 1915: 456 as Chironomus) - IL, widespread

      Micropsectra Kieffer, 1909: 50 (Tribe Tanytarsini)

        Micropsectra connexa (Kieffer, 1906: 17 as Chironomus) - ON, widespread north
          (syn.) Chironomus brunneus Walker, 1848: 21
        Micropsectra dives (Johannsen, 1905: 288 as Tanytarsus) - OH, WI
        Micropsectra geminata Oliver and Dillon 1994 - Egan & Ferrington 2015: 16 (Isle Royale, pupal exuviae)
        Micropsectra logani (Johannsen, 1928: 33 as Tanytarsus) - Cook-MSU, Egan & Ferrington 2015: 16 (Isle Royale, pupal exuviae)
        Micropsectra nigripila (Johannsen, 1905: 287 as Tanytarsus) - Cook-MSU
        Micropsectra polita (Malloch, 1915: 493 as Tanytarsus) - IL, widespread
        Micropsectra recurvata Goetghebuer, 1928: 117 - widespread east
        Micropsectra subletteorum Anderson, Stur and Ekrem 2013: 906 - Egan & Ferrington 2015: 16 (Isle Royale, pupal exuviae)
        Micropsectra xantha (Roback, 1955: 4 as Calopsectra(Micropsectra)) - Hudson & Bright 2012, Egan & Ferrington 2015: 15 (Isle Royale, pupal exuviae)

      Microtendipes Kieffer, 1915: 70 (Tribe Chironomini)

        Microtendipes caducus Townes, 1945: 24 - Townes 1945
        Microtendipes caelum Townes, 1945: 23 - OH, ON, northeast
        Microtendipes pedellus (De Geer, 1776: 378 as Tipula) - Townes 1945
          (syn.) Chironomus fascipes Coquillett, 1908: 145
          (syn.) Chironomus deflexa Walley, 1926b: 206

      Neostempellina Reiss, 1984: 204 (Tribe Tanytarsini)

        Neostempellina reissi Caldwell, 2000: 164 - OH (expect in s. MI - sand substrates of springruns and 1st order streams)

      Nilothauma Kieffer, 1921b: 270 (Tribe Chironomini)

        Nilothauma babiyi (Rempel, 1937b: 274 as Chironomus) - Townes 1945
        Nilothauma bicorne (Townes, 1945: 35 as Kribioxenus) - Cook-MSU
        Nilothauma mirabile (Townes, 1945: 35 as Kribioxenus) - Hudson & Bright 2012

      Omisus Townes, 1945: 27 (Tribe Chironomini)

        Omisus pica Townes, 1945: 27 - widespread

      Pagastiella Brundin, 1949: 840 (Tribe Chironomini)

        Pagastiella ostansa (Webb, 1969: 102 as Paralauterborniella) - ON, widespread
          unassociated larvae of likely this species has been collected from Lake Huron by P. Hudson

      Parachironomus Lenz, 1921: 160 (Tribe Chironomini)

        Parachironomus abortivus (Malloch, 1915 as Chironomus) - IL, ON, widespread
        Parachironomus carinatus (Townes, 1945 as Harnischia) - OH, widespread
        Parachironomus forceps (Townes, 1945 as Harnischia) - Townes 1945
        Parachironomus frequens (Johannsen, 1905 as Chironomus) - Townes 1945
        Parachironomus hazelriggi Spies, 2000: 133 - OH, widespread, principally south
        Parachironomus hirtalatus (Beck & Beck, 1964 as Chironomus) - IL, east
        Parachironomus monochromus (Wulp, 1874: 129 as Chironomus) - Oliver et al. 1990
        Parachironomus pectinatellae (Dendy & Sublette, 1959 as Tendipes) - IL, east
        Parachironomus potamogeti (Townes, 1945 as Harnischia) - Cook-MSU
        Parachironomus tenuicaudatus (Malloch, 1915 as Chironomus) - Cook-MSU
        Parachironomus varus (Goetghebuer, 1921: 162 as Chironomus) - widespread north

      Paracladopelma Harnisch, 1923: 304 (Tribe Chironomini)

        Paracladopelma galaptera (Townes, 1945: 148 as Harnischia) - Townes 1945
        Paracladopelma loganae Beck & Beck, 1969: 303 - KS-OH, s. to MS-FL
        Paracladopelma nais (Townes, 1945: 149 as Harnischia) - Townes 1945
        Paracladopelma nereis (Townes, 1945: 150 as Harnischia) - Townes 1945
        Paracladopelma undine (Townes, 1945: 149 as Harnischia) - Cook-MSU
        Paracladopelma winnelli Jackson, 1977: 1334 - Jackson 1977

      Paralauterborniella Lenz, 1941: 48 (Tribe Chironomini)

        Paralauterborniella nigrohalteralis (Malloch, 1915: 440 as Chironomus) - Townes 1945

      Paratanytarsus Thienemann & Bause, 1913: 120 (Tribe Tanytarsini)

        Paratanytarsus dimorphis Reiss, 1965: 496 as Tanytarsus) - Egan & Ferrington 2015: 17 (Isle Royale, pupal exuviae)
        Paratanytarsus dissimilis (Johannsen, 1905: 292 as Tanytarsus) - ON, widespread
          (syn.) Paratanytarsus confusus Palmén, 1960: 283
        Paratanytarsus dubius (Malloch, 1915: 496 as Tanytarsus) - IL
        Paratanytarsus grimmii (Schneider, 1885: 301 as Chironomus) - ON, widespread north
          (syn.) Tanytarsus boiemicus Bause, 1913: 97
        Paratanytarsus inopertus (Walker, 1856: 164 as Chironomus) - WI
        Paratanytarsus laccophilus (Edwards, 1929: 409 as Tanytarsus) - Egan & Ferrington 2015: 16 (Isle Royale, pupal exuviae)
        Paratanytarsus longistilus Bolton, Ekrem, Sublette and Sublette 2010: 264 - Washtenaw Co., Fleming Cr., Cherry Hill Rd., 20-VII-1988, M. J. Bolton
        Paratanytarsus natvigi Goetghebuer, 1933: 21 - Egan & Ferrington 2015: 17 (Isle Royale, pupal exuviae)
        Paratanytarsus similatus (Malloch, 1915: 494 as Tanytarsus) - WI
        Paratanytarsus sp. nov. (Bolton et al.) - WI (expect in MI in cool springs)

      Paratendipes Kieffer, 1911a: 41 (Tribe Chironomini)

        Paratendipes albimanus (Meigen, 1818: 40 as Chironomus) - Townes 1945
        Paratendipes basidens Townes, 1945: 30 - widespread
        Paratendipes duplicatus (Johannsen, 1937b: 28 as Chironomus) - Townes 1945
        Paratendipes nitidulus (Coquillett, 1901a: 608 as Chironomus) - Oliver et al. 1990: 50
        Paratendipes subaequalis (Malloch, 1915: 440 as Chironomus) - OH, widespread

      Phaenopsectra Kieffer, 1921b: 274 (Tribe Chironomini)

        Phaenopsectra dyari (Townes, 1945: 75 as Tanytarsus) - OH, widespread
        Phaenopsectra flavipes (Meigen, 1818: 50 as Chironomus) - Cook-MSU
        Phaenopsectra obediens (Johannsen, 1905: 286 as Tanytarsus) - UMMZ; Cook-MSU
        Phaenopsectra punctipes (Wiedemann, 1817: 65 as Chironomus) - Cook-MSU

      Polypedilum Kieffer, 1912b: 12 (Tribe Chironomini)

        Polypedilum (Tripodura) acifer Townes, 1945 - Townes 1945
        Polypedilum (Polypedilum) albicorne (Meigen, 1838: 6 as Chironomus) - Cook-MSU
          (syn.) Polypedilum nigritum Townes, 1945: 56
        Polypedilum (Tripodura) albinodus Townes, 1945 - IL, OH, ON, widespread
        Polypedilum (Polypedilum) angustum Townes 1945: 56 - Hudson & Bright 2012
        Polypedilum (Polypedilum) artifer (Curran, 1930: 32 as Chironomus) - Cook-MSU
        Polypedilum (Polypedilum) atrichon Maschwitz 2000: 43 - MN, ON
        Polypedilum (Uresipedilum) aviceps Townes, 1945: 61 - OH, ON, widespread
        Polypedilum (Polypedilum) bergi Maschwitz 2000: 49 - Maschwitz & Cook 2000
        Polypedilum (Polypedilum) braseniae (Leathers, 1922: 23 as Chironomus) - Cook-MSU; Otsego Co., Perch Lake - EB pers. obs.)
        Polypedilum (Polypedilum) calopterum (Mitchell, 1908: 8 as Chironomus) - Townes 1945
          (syn.) Polypedilum walleyi Townes, 1945: 52, Maschwitz 2000: 57
        Polypedilum (Polypedilum) cinctum Townes, 1945: 59 - widespread middle latitudes
        Polypedilum (Uresipedilum) convictum (Walker, 1856a: 161 as Chironomus) - Cook-MSU
        Polypedilum (Tripodura) digitifer Townes, 1945 - Cook-MSU
        Polypedilum (Polypedilum) fallax (Johannsen, 1905: 210 as Chironomus) - Townes 1945
        Polypedilum (Polypedilum) flavum (Johannsen, 1905: 225 as Chironomus) - Townes 1945
          (syn.) Chironomus flaviventris Johannsen, 1907: 111
          (syn.) Polypedilum obtusum Townes, 1945: 59, in part
        Polypedilum (Tripodura) gomphus Townes, 1945 - ON, widespread
        Polypedilum (Polypedilum) griseopunctatum (Malloch, 1915) - OH
        Polypedilum (Tripodura) halterale (Coquillett, 1901b as Chironomus) - Townes 1945
          (syn.) Chironomus griseopunctatus Malloch, 1915
          (syn.) Polypedilum digitifer Townes, 1945
          (syn.) Polypedilum simulans Townes, 1945
        Polypedilum (Polypedilum) illinoense (Malloch, 1915: 471 as Chironomus) - Townes 1945; Berg 1950: 91-92
        Polypedilum (Polypedilum) laetum (Meigen, 1818: 30 as Chironomus) - Yanoviak & McCafferty 1996 (Mountain Stream, Salmon-Trout, Pine and Huron Rivers, Marquette Co.)
          (syn.) Chironomus nubilipennis Meigen, 1838: 3 (fide Goetghebuer 1923: 127), Gimmerthal, 1845: 319
          (syn.) Tendipes falciger (Kieffer), Kieffer, 1911: 21 (fide Goetghebuer 1923: 126), Ashe and Cranston, 1991: 132
          (nom.dub.) Polypedilum heptastictum Kieffer, 1922: 258
        Polypedilum (Polypedilum) nubeculosum (Meigen, 1804: 18 as Chironomus) - Cook-MSU
          (syn.) Chironomus enotatus Walker, 1856: 170 (fide Goetghebuer 1928: 89), Kieffer 1906: 17
          (syn.) Polypedilum emarginatum Kieffer, 1913: 16 (fide Edwards 1929: 402), Kieffer 1920: 161
          (nom. dub.) Polypedilum hirtimanus Kieffer, 1915: 82
          (nom. dub.) Polypedilum ciliatimanus Kieffer, 1915: 82
          (nom. dub.) Polypedilum obliteratum Kieffer, 1920: 164, var. opacum Kieffer, 1922b: 92, var. majus Kieffer, 1922b: 91
        Polypedilum (Polypedilum) newaygo Taber, 2009 - Taber 2009
        Polypedilum (Polypedilum) nymphaeorum Maschwitz 2000: 92 - Maschwitz & Cook 2000
        Polypedilum (Uresipedilum) obtusum Townes, 1945: 60 - Cook-MSU
        Polypedilum (Polypedilum) ontario (Walley, 1926b: 206 as Chironomus) - OH, ON, widespread
          (syn.) Chironomus hirtipes Mitchell, 1908
        Polypedilum (Polypedilum) ophioides Townes, 1945: 57 - Townes 1945; Berg 1950: 92
        Polypedilum (Polypedilum) pedatum Townes, 1945: 55 - Townes 1945
        Polypedilum (Polypedilum) pedestre (Meigen, 1830: 246 as Chironomus) - widespread
          (nom. dub., syn.?) Chironomus fuscipennis Meigen, 1818: 35
          (syn.) Chironomus pictipes Zetterstedt, 1850: 3569 (fide Edwards 1929: 400)
          (syn.) Chironomus patens Walker, 1856: 170 (fide Goetghebuer 1928: 91)
        Polypedilum (Tripodura) scalaenum (Schrank, 1803: 73 as Tipula) - Townes 1945
          (syn.) Chironomus needhamii Johannsen, 1908: 278
        Polypedilum (Pentapedilum) sordens (van der Wulp, 1874: 141 as Tanytarsus) - Townes 1945; Berg 1950: 96-97
          (syn.) Tanytarsus fulvescens Johannsen, 1905: 293
        Polypedilum (Pentapedilum) tritum (Walker, 1856a: 342 as Chironomus) - Hudson & Bright 2012
          (syn.) Polypedilum albulum Townes, 1945: 63
          (syn.) Pentapedilum fulvescens? Townes, 1937: 171
        Polypedilum (Polypedilum) trigonus Townes, 1945: 49 - Townes 1945
        Polypedilum (Polypedilum) tuberculum Maschwitz 2000: 119 - MN, WI
        Polypedilum (Polypedilum) vibex Townes, 1945: 52 - ON, widespread
        Polypedilum (Polypedilum) walleyi Townes, 1945: 52 - ON
          (syn.) Chironomus calopterum Mitchell, 1908, Boesel 1985: 257, Maschwitz 2000: 124

      Pseudochironomus Malloch, 1915 (Subfamily Pseudochironomini)

        Pseudochironomus anas Townes, 1945: 17 - Cook-MSU
        Pseudochironomus articaudus Sæther, 1977: 81 - ON, widespread north
        Pseudochironomus banski Townes, 1945: 17 - Cook-MSU
        Pseudochironomus chen Townes, 1945: 18 - ON, widespread
        Pseudochironomus fulviventris (Johannsen, 1905: 229 as Chironomus) - Townes 1945: 16
        Pseudochironomus julia (Curran, 1930: 34 as Orthocladius) - Townes 1945: 19, Oliver et al. 1990: 55
          (syn.) Pseudochironomus aix Townes, 1945: 19
        Pseudochironomus middledauffi Townes, 1945: 18 - Cook-MSU
        Pseudochironomus netta Townes 1945: 19 - Townes 1945: 19
        Pseudochironomus pseudoviridis (Malloch, 1915: 450 as Chironomus) - Cook-MSU
        Pseudochironomus rex Hauber, 1947: 458 - MB-FL, OH, widespread
        Pseudochironomus richardsoni Malloch, 1915 - OH, ON, widespread

      Rheotanytarsus Thienemann & Bause, 1913: 120 (as subgenus of Syntanytarsus) (Tribe Tanytarsini)

        [Note: Undergoing revision as Torbjørn Ekrem and Lisa Stur (based on Sublette material) describe new species from North America]
        Rheotanytarsus exiguus (Johannsen, 1905: 294 as Tanytarsus) - (EB, pers. obs.)
        Rheotanytarsus pellucidus (Walker, 1848: 21 as Chironomus) - (EB, pers. obs.)
          (syn.) Stempellinella distinctissimus Brundin, 1947: 88
          (syn.) Rheotanytarsus distinctissimus (Brundin, 1947: 88), Lehmann 1970

      Robackia Sæther, 1977: 123 (Tribe Chironomini)

        Robackia claviger (Townes, 1945: 158 as Harnischia) - IN, widespread
        Robackia demeijerei (Kruseman, 1933: 195 as Tendipes) - Winnell and White 1985
        Robackia pilicauda Sæther, 1977: 129 - Oliver & Dillon, 1994

      Saetheria Jackson, 1977: 1325 (Tribe Chironomini)

        Saetheria tylus (Townes, 1945: 150 as Harnischia) - Winnell and White 1985

      Sergentia Kieffer, 1922a: 288 (Tribe Chironomini)

        Sergentia coracina (Zetterstedt, 1850: 3508 as Chironomus) - Egan & Ferrington 2015: 17 (Isle Royale, pupal exuviae)

      Stelechomyia Reiss, 1982: 294 (Tribe Chironomini)

        Stelechomyia perpulchra (Mitchell, 1908: 13 as Chironomus) - UMMZ (larvae live on submersed wood)

      Stempellina Thienemann & Bause, 1913: 120 (Tribe Tanytarsini)

        [note: larvae of the very distinctive Stempellina have been collected from several sites in Michigan, although none yet have been associated to a species]
        Stempellina ranota Webb, 1969: 96 - ON
        Stempellina rodesta Webb, 1969: 95 - ON

      Stempellinella Brundin, 1947: 87 (Tribe Tanytarsini)

        Stempellinella boltoni Ekrem, 2007: 1378 - OH
        Stempellinella edwardsi Spies & Sæther, 2004: 65 - Ekrem 2007: 1405
          (syn.) Tanytarsus (Stempellina) minor Edwards, 1929: 420
          (syn.) Tanytarsus (Stempellina) minusculus Edwards, 1929: 417
        Stempellinella fimbriata Ekrem, 2007: 1411 - Ekrem 2007
          Ekrem (2007) indicates that all NA specimens identified as S. brevis are S. fimbriata]
        Stempellinella leptocelloides (Webb, 1969: 94 as Stempellina) - MN, OH

      Stenochironomus Kieffer, in Kieffer and Thienemann, 1919: 44 (Tribe Chironomini)

        [larvae are wood miners, pupae found in the longitudinal chambers made by larvae]
        Stenochironomus cinctus Townes, 1945: 87 - Hudson & Bright 2012
          (syn.) Stenochironomus aestivalis Townes, 1945: 86
        Stenochironomus (Stenochironomus) hilaris (Walker, 1848: 17 as Chironomus) - Townes 1945
          (syn.) Chironomus tæniapennis Coquillett, 1901a: 607-608
          (syn.) Chironomus nephopterus Mitchell, 1908: 7
          (syn.) Chironomus exquistus Mitchell, 1908: 11
          (syn.) Chironomus zonopterus Mitchell, 1908: 12
        Stenochironomus (Stenochironomus) macateei (Malloch, 1915: 45 as Chironomus) - Hudson & Bright 2012
        Stenochironomus (Stenochironomus) poecilopterus (Mitchell, 1908: 10 as Chironomus) - Cook-MSU
        Stenochironomus (Stenochironomus) pulchripennis (Coquillett, 1902b: 94 as Chironomus) - Oliver et al. 1990:53

      Stictochironomus Kieffer, in Kieffer and Thienemann, 1919: 44 (Tribe Chironomini)

        Stictochironomus annulicrus (Townes, 1945: 83 as Tanytarsus) - OH, ON, widespread
        Stictochironomus devinctus (Say, 1829: 150 as Chironomus) - Townes 1945
          (syn.) Chironomus compes Coquillett, 1908: 145
          (syn.) Chironomus ornatipes Kieffer, 1917: 343
        Stictochironomus marmoreus (Townes, 1945: 81 as Tanytarsus) - Cook-MSU
        Stictochironomus palliatus (Coquillett, 1902b: 95 as Chironomus) - WI, widespread e. 1/2 temperate NA
        Stictochironomus varius (Townes, 1945: 82 as Tanytarsus) - Townes 1945

      Tanytarsus Wulp, 1874: 134 (Tribe Tanytarsini)

        [note: this genus is undergoing a thorough review by Dr. Ekrem and others; a recent review of several species groups has recently been published]
        Tanytarsus acifer Ekrem, Sublette & Sublette, 2003: 266 - Ekrem et al. 2003: 271
          (syn.) Tanytarsus n. sp. Ruse et al. 2000: 56
        Tanytarsus aigos Ekrem, Sublette & Sublette, 2003: 275 - Ekrem et al. 2003: 276
        Tanytarsus nr. angulatus Kawai, 1991: 168 - OH, widespread (see note in Ekrem et al. 2003: 279-280)
        Tanytarsus boltoni Ekrem, Sublette & Sublette, 2003: 284 - OH
        Tanytarsus brundini Lindeberg, 1963: 127 - Hudson & Bright 2012
        Tanytarsus buckleyi Sublette, 1964b: 143 - Hudson & Bright 2012
        Tanytarsus confusus Malloch, 1915: 490 - Ekrem et al. 2003: 287
        Tanytarsus dendyi Sublette, 1964b: 141 - OH, widespread south, maybe s. LP
        Tanytarsus dispar Lindeberg, 1967: 55 - widespread north, Holarctic, may be synonym of Tanytarsus lestagei (see note in Ekrem et al. 2003, p, 305)
        Tanytarsus epleri Ekrem, Sublette & Sublette, 2003: 291 - OH (Lake Erie), widespread SE, expect in southeastern LP
        Tanytarsus fatigans Johannsen, 1905: 292 - widespread
        Tanytarsus glabrescens Edwards, 1929: 436 - IL
        Tanytarsus guerlus (Roback, 1957a: 133 as Calopsectra) - IL, OH, MN, widespread
        Tanytarsus herrmanni Ekrem, Sublette & Sublette, 2003: 300 - CO, NM, ON, widespread
        Tanytarsus inaequalis Goetghebuer, 1921: 118 - Hudson & Bright 2012
        Tanytarsus lestagei Goetghebuer, 1922: 47 - OH, widespread
        Tanytarsus lobiger Ekrem, Sublette & Sublette, 2003: 305 - Ekrem et al. 2003: 305
        Tanytarsus martini Ekrem, Sublette & Sublette, 2003: 306 - Ekrem et al. 2003: 306
        Tanytarsus mendax Kieffer, 1925b: 223 - MN, OH, ON, widespread
          (syn.) Tanytarsus holochlorus Edwards, 1929: 414
          (syn.) Tanytarsus xanthus Sublette, 1964b: 142
        Tanytarsus messersmithi Ekrem, Sublette & Sublette, 2003: 310 - MN, OH, widespread
        Tanytarsus muticus Johannsen, 1905: 294 - IL, OH, WI
        Tanytarsus neoflavellus Malloch, 1915: 489 - Ekrem et al. 2003: 313
        Tanytarsus occultus Brundin, 1949: 847 - WI, mainly west Palearctic but also northeastern Nearctic
        Tanytarsus seppi Ekrem, Sublette & Sublette, 2003: 317 - OH, ON, widespread
        Tanytarsus volgensis Miseiko, 1967: 55 - Palearctic and Nearctic, widespread
        Tanytarsus wirthi Ekrem, Sublette & Sublette, 2003: 320 - Egan & Ferrington 2015: 17 (Isle Royale, pupal exuviae)

      Tribelos Townes, 1945: 66 (Tribe Chironomini)

        Tribelos atrum (Townes, 1945: 68 as Tanytarsus) - Grodhaus 1987
        Tribelos fuscicorne (Malloch, 1915: 466 as Tanytarsus) - Townes 1945
        Tribelos jucundus (Walker, 1848: 16 as Tanytarsus) - UMMZ
          (syn.) Chironomus dimorphus Malloch, 1915: 464
          (syn.) Chironomus fusciventris Malloch, 1915: 465
          (syn.) Tanytarsus protextus Townes, 1945 : 69

      Xenochironomus Kieffer, 1921b: 273 (Tribe Chironomini)

        Xenochironomus xenolabis (Kieffer, in Thienemann & Kieffer, 1916: 526 as Chironomus) - Pat Hudson, pers. comm. 2011

      Zavrelia Kieffer, Thienemann & Bause in Bause 1913: 73 (Tribe Tanytarsini)

        Zavrelia aristata Ekrem & Stur, 2009: 121 - OH (Ekrem and Stur 2009: 125)

      Zavreliella Kieffer, 1920b: 334 (Tribe Chironomini)

        Zavreliella varipennis (Coquillett, 1902b: 94 as Chironomus) - Oliver et al. 1990:55

    Subfamily DIAMESINAE

      Boreoheptagyia Brundin, 1966: 420 (Tribe Boreoheptagyiini)

        Boreoheptagyia lurida (Garrett, 1925a: 6) Diamesa (orig.) - Northern Nearctic, expect in UP

      Diamesa Meigen, in Gistl 1835: 66 (Tribe Diamesini)

        Diamesa chiobates Hansen, in Hansen & Cook, 1976:70 - MN, WI (Oliver et al. 1990:17)
        Diamesa davisi Edwards, 1933: 614 - NWT, UT, NH (Oliver et al. 1990:17)
        Diamesa incallida (Walker, 1856a: 183 as Chironomus) - ON, widespread northern Holarctic (expect in UP)
          (syn.) Chironomus nexilis Walker, 1856
          (syn.) Diamesa fonticola Sæther, 1969: 24
          (syn.) Diamesa sp. VII Thienemann, 1941
          (syn.) Psilodiamesa incallida (Walker), 1856: 183
        Diamesa insignipes Kieffer, in Kieffer & Thienemann, 1908: 3 - Egan & Ferrington 2015: 4 (Isle Royale, pupal exuviae)
        Diamesa leona Roback, 1957b - WA-MN, QUE, NB
          (syn.) Diamesa caena Roback, 1957b
          (syn.) Diamesa pieta Roback, 1957b
        Diamesa mendotae Muttkowski, 1915: 116 - MN, WI
        Diamesa nivoriunda (Fitch, 1847: 282 as Chironomus) - Hanson and Cook 1976

      Lappodiamesa Serra-Tosio, 1968: 140 (Tribe Diamesini)

        Lappodiamesa boltoni Sæther & Willassen, 1988: 78 - OH

      Pagastia Oliver, 1959: 49 (Tribe Diamesini)

        Pagastia orthogonia Oliver, 1959: 51 - Oliver and Roussel 1982, Caldwell 2007: 69, Egan & Ferrington 2015: 4 (Isle Royale, pupal exuviae)

      Potthastia Kieffer, 1922c: 362 (Tribe Diamesini)

        Potthastia gaedii (Meigen, 1838) Diamesa (orig.) - e. USA (an adult specimen near gaedii was collected in Lake Superior by P. Hudson (P. Hudson, pers. comm. Dec. 2009), Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae)
        Potthastia iberica Serra-Tosio, 1971: 160 - WY, GA
        Potthastia longimana Kieffer, 1922c: 362 - Winnell and White 1985

      Protanypus Kieffer, 1906: 40 (Tribe Protanypini)

        Protanypus hamiltoni Sæther 1975d: 375 - Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae)
        Protanypus ramosus Sæther 1975d: 368- adult from Lake Superior, larvae from Lakes Michigan and Huron (P. Hudson, pers. comm. Dec. 2009), Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae)

      Pseudodiamesa Goetghebuer, 1939: 9 (Tribe Diamesini)

        [note: larvae of an unplaced species of Pseudodiamesa were collected in Marquette Co. from a spring-fed brook in the Huron Moutains by EB, 1998]
        Pseudodiamesa (Pseudodiamesa) branickii (Nowicki, 1873: 3 as Diamesa) - Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae), expect widespread in UP

      Sympotthastia Pagast, 1947: 547 (Tribe Diamesini)

        Sympotthastia n. sp. - adult of an undescribed species from Lake Huron (P. Hudson, pers. comm. Dec. 2009)

    Subfamily ORTHOCLADIINAE

      Acamptocladius Brundin, 1956: 162

        Acamptocladius dentolatens (Sæther, 1971: 1814 as Phycoidella) - ON

      Acricotopus Kieffer, 1921a: 90

        Acricoptus nitidellus (Malloch, 1915: 515 as Orthocladius) - BC-MB, IL, NY
          (syn.) Spaniotoma senex Johannsen, 1937a: 63

      Allocladius Kieffer, 1913d: 28

        Allocladius arenarius (Strenzke, 1960: 414 as Psuedosmittia) - widespread
          (syn.) Pseudosmittia gracilis Thienemann & Strenzke, 1940: 33
        Allocladius nanseni (Kieffer, 1926: 85 as Camptochironomus) - Ferrington and Saether 2011
          (syn.) Pseudosmittia mediocarinata Caspers et Reiss, 1989: 132
          (syn.) Smittia longitibia Goetghebuer, 1933: 29

      Baeoctenus Sæther, 1976: 41

        Baeoctenus bicolor Sæther, 1976: 43 - MB, NB (associated with freshwater clam Anodonta)

      Brilla Kieffer, 1913b: 34

        Brilla flavifrons (Johannsen, 1905: 301 as Metriocnemus) - Oliver 1983
        Brillia parva Johannsen, 1934: 351 - ON (e. UP record possible)

      Bryophaenocladius Thienemann, 1934: 36

        Bryophaenocladius digitatus Sæther, 1973a: 55 - SD, NC, SC
        Bryophaenocladius flavoscutellatus (Malloch, 1915: 523 as Orthocladius) - IL, ON east
        Bryophaenocladius pleuralis (Malloch, 1915: 527 as Orthocladius) - IL
        Bryophaenocladius subparallelus (Malloch, 1915: 522 as Orthocladius) - SD, IL, OH

      Camptocladius Wulp, 1874: 133

        Camptocladius stercorarius (De Geer, 1776: 388 as Tipula) - widespread northeast
          (syn.) Tipula byssina Shrank, 1803: 76

      Cardiocladius Kieffer, 1912a: 22

        Cardiocladius albiplumus Sæther, 1969:34-36 - Bode 1983 (as Eukiefferiella similis gp.)
        Cardiocladius fulvus (Johannsen, 1908: 275 as Thalassomyia) - Oliver et al. 1990: 21
        Cardiocladius obscurus (Johannsen, 1903: 437 as Thalassomyia) - OH, ON, widespread
          (nom. nud.) Thalassomyia fusca Johannsen, 1903: 437

      Chaetocladius Kieffer, 1911b: 182

        Chaetocladius piger (Goetghebuer, 1913: 157 as Dactylocladius) - OH, ON, widespread (Caldwell 1997)
          (syn.) Chaetocladius stamfordi (Johannsen, 1947: 171 as Hydrobaenus)
          (syn.) Chaetocladius oliveri Sæther, 1969: 93

      Chasmatonotus Loew, 1864: 51

        Chasmatonotus atripes Rempel, 1937a: 253 - UMMZ
        Chasmatonotus unimaculatus Loew, 1864: 50 - Oliver et al. 1990:22

      Corynoneura Winnertz, 1846: 12

        Corynoneura arctica Kieffer, 1923: 4 - Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae)
        Corynoneura celeripes Winnertz, 1852: 50 - IL, IN, OH, ON
          (syn.) Corynoneura atra Winnertz, 1852: 50
        Corynoneura doriceni Makarchenko and Makarchenko, 2006: 12 - Egan & Ferrington 2015: 6 (Isle Royale, pupal exuviae)
        Corynoneura lobata Edwards, 1924: 186 - pers. obs. (EB)
        Corynoneura oxfordana Boesel & Winner, 1980: 505 - Hudson & Bright 2012
        Corynoneura scutellata Winnertz, 1846: 13 - widespread

      Cricotopus Wulp, 1874: 132

        Cricotopus (Cricotopus) absurdus Johannsen, 1905: 277 - OH, widespread
        Cricotopus (Cricotopus) annulator Goetghebuer, 1927: 52 - Egan & Ferrington 2015: 6 (Isle Royale, pupal exuviae)
          (syn.) Cricotopus irwini Sublette & Sublette, 1971: 97
        Cricotopus (Cricotopus) bicinctus (Meigen, 1818: 41 as Chironomus) - UMMZ, Egan & Ferrington 2015: 6 (Isle Royale, pupal exuviae)
        Cricotopus (Cricotopus) brunnicans Walley, 1928a: 21 - Cook-MSU
        Cricotopus (Cricotopus) coronatus Hirvenoja, 1973: 222 - widespread north
        Cricotpus (Cricotopus) curtus Hirvenoja, 1973: 199 - Egan & Ferrington 2015: 6 (Isle Royale, pupal exuviae)
        Cricotopus (unassigned) diversus Boesel, 1983: 85 - Oliver et al. 1990:25
        Cricotopus (Cricotopus) elegans Johannsen, 1943: 78 - Johannsen, 1943; Berg 1950: 88-90
        Cricotopus (Cricotopus) festivellus (Kieffer, 1906: 18 as Chironomus) - ON, OH
        Cricotopus (Isocladius) flavibasis Malloch, 1915: 502 - IL west (expect in w. UP)
        Cricotopus (Isocladius) flavipes Johannsen, 1942: 73 - Johannsen, 1942; Berg 1950: 90-91
        Cricotopus (Cricotopus) fuscus (Kieffer, 1909: 46 as Trichocladius) - Egan & Ferrington 2015: 6-7 (Isle Royale, pupal exuviae)
        Cricotopus (Cricotopus) infuscatus Malloch, 1915: 517 - IL, OH, ON, widespread
          (syn.) Cricotopus aratus Roback, 1957a: 73
          (syn.) Cricotopus ceris Roback, 1957a: 72
        Cricotopus (Isocladius) intersectus (Staeger, 1839: 574) - Chironomus (orig.) - Egan & Ferrington 2015: 7 (Isle Royale, pupal exuviae)
        Cricotopus (Cricotopus) luciae LeSage in LeSage & Harrison: 1980: 91 - OH, ON
        Cricotopus (Isocladius) myriophylli Oliver, 1984: 1287 - ON (expect in northern MI associated with Myriophyllum spicatum)
        Cricotopus (Nostococladius) nostococicola Wirth, 1957: 122 - OH, widespread
        Cricotopus (Paratrichocladius) nitidus (Malloch, 1915: 515 as Orthocladius) - IL, widespread north and west
          (syn.) Trichocladius helis Roback, 1957b: 15
          (syn.) Syncricotopus fontinalis Sæther, 1969: 77
        Cricotopus (unassigned) olivetus Boesel, 1983: 88 - OH (in Boesel 1983)
        Cricotopus (Isocladius) ornatus (Meigen, 1818: 43 as Chironomus) - ON, widespread north
        Cricotopus (Isocladius) pilitarsis (Zetterstedt, 1850: 3565 as Chironomus) - widespread north
        Cricotopus (Cricotopus) politus (Coquillett, 1902b: 93 as Orthocladius) - IL, OH, widespread
        Cricotopus (Paratrichocladius) rufiventris (Meigen, 1830: 249 as Chironomus) - ON, OH
        Cricotopus (Cricotopus) slossonae Malloch, 1915: 506 - IL, OH, ON
        Cricotopus (Isocladius) sylvestris (Fabricius, 1794: 252 as Tipula) - Egan & Ferrington 2015: 7 (Isle Royale, pupal exuviae)
        Cricotopus (Cricotopus) tremulus (Linneaus, 1758: 587 as Tipula) - Egan & Ferrington 2015: 7 (Isle Royale, pupal exuviae)
        Cricotopus (Cricotopus) triannulatus (Macquart, 1826: 202 as Chironomus) - Egan & Ferrington 2015: 7 (Isle Royale, pupal exuviae)
        Cricotopus (Isocladius) tricinctus (Meigen, 1818: 41 as Chironomus) - OH, widespread
          (syn.) Cricotopus lebetis Sublette, 1964b: 118
        Cricotopus (Cricotopus) trifascia Edwards, 1929: 322 - SK-PA, OH
          (syn.) Cricotopus ithacanensis Sublette, 1967b: 555
        Cricotopus (Cricotopus) tristus Hirvenoja 1973: 196 - Egan & Ferrington 2015: 7 (Isle Royale, pupal exuviae)
        Cricotopus (Isocladius) trifasciata (Meigen, in Panzer, 1813 as Chironomus) - Cook-MSU; Berg 1950: 91
          (syn.) Cricotopus remus Sublette, 1964b: 115
        Cricotopus (Cricotopus) varipes Coquillett, 1902: 93 - UMMZ
        Cricotopus (Cricotopus) vierriensis Goetghebuer, 1935: 417 - OH widespread

      Diplocladius Kieffer, in Kieffer & Thienemann, 1908: 6

        Diplocladius cultriger Kieffer, in Kieffer & Thienemann 1908: 6 - OH, widespread

      Diplosmittia Sæther, 1981b: 29

        Diplosmittia carinata Sæther, 1985b: 55 - Sæther, 1985

      Doithrix Sæther & Sublette, 1983: 6

        Doithrix villosa Sæther & Sublette, 1983: 9 - widespread [note: probably semiterrestrial, found in floodplains near margins of of small streams and seepage areas]

      Doncricotopus Sæther, 1981a: 223

        Doncricotopus sp. - larvae collected from from unspecified location in Lake Superior (Stroom et al. 2003)

      Epoicocladius Zavrel, in Šulc & Zavrel 1924: 368

        [note: larvae are epizootic or phoretic on bodies of the larval ephemerid mayflies (Ephemeroptera: Ephemeridae)]
        Epoicocladius flavens (Malloch, 1915) Camptocladius (orig.) - Oliver et al. 1990:25 (associated with Hexagenia spp.)
          (syn.) Camptocladius ephemerae Kieffer, in Sulc & Zavrel 1924
        Epoicocladius sp. 2 - associated with Litobranchia recurvata (Rick Jacobsen, pers. comm. 2004)
        Epoicocladius sp. 3 - associated with Ephemera spp. (Rick Jacobsen, pers. comm. 2004)

      Eukiefferiella Thienemann, 1926b: 325

        Eukiefferiella brevinervis (Malloch, 1915 as Orthocladius) - IL, OH, PA
        Eukiefferiella claripennis (Lundbeck, 1898: 281 as Chironomus) - Egan & Ferrington 2015: 8 (Isle Royale, pupal exuviae)
        [note: larvae of this species or very close have been sampled in streams in Michigan - EB, pers. obs.]
          (syn.) Spaniotoma hospita Edwards, 1929: 351
        Eukiefferiella coerulescens (Kieffer, 1926 as Trichocladius) - Egan & Ferrington 2015: 8 (Isle Royale, pupal exuviae)
        Eukiefferiella devonica (Edwards, 1929: 349 as Spaniotoma) - widespread
        [note: larvae of this species or very close have been sampled in streams in Michigan - EB, pers. obs.]

      Gymnometriocnemus Goetghebuer, 1932: 23

        Gymnometriocnemus (Raphidocladius) brumalis (Edwards, 1929: 316 as Metriocnemus) - widespread
          (syn.) Gymnometriocnemus acigus Sæther, 1983a: 214
        Gymnometriocnemus (Gymnometriocnemus) marionensis Sæther, 1969: 129 - Stur and Ekrem 2015
        Gymnometriocnemus (Gymnometriocnemus) subnudus (Edwards, 1929: 316 as Metriocnemus) - ON, widespread north and east

      Heleniella Gowin, 1943: 116

        Heleniella hirta Sæther, 1969: 134 - Pat Hudson (pers. comm., 2012)
        Heleniella sp. Sæther, 1969 - male of an unidentified species collected from w. Washtenaw Co. by Pat Hudson

      Heterotrissocladius Spãrck, 1923: 94

        Heterotrissocladius changi Sæther, 1975a: 47 - Winnell and White 1985; Egan & Ferrington 2015: 8 (Isle Royale, pupal exuviae)
        Heterotrissocladius cooki Sæther, 1975: 41 - MN
        Heterotrissocladius marcidus (Walker, 1856a: 177 as Chironomus) - Sæther, 1975
        Heterotrissocladius oliveri Sæther, 1975: 19 - Sæther, 1975 (indicated as MN but off Isle Royale, 27.5-120.5m depth in Lake Superior: p. 22); Egan & Ferrington 2015: 8 (Isle Royale, pupal exuviae)

      Heterotanytarsus Spärck, 1923: 92

        Heterotanytarsus nudalus Sæther, 1975c: 263 - ON, expect in northern oligotrophic lakes

      Hydrobaenus Fries, 1830: 177

        Hydrobaenus johannseni (Sublette, 1967b: 504 as Orthocladius) - Hudson coll.
          (syn.) Orthocladius domus Sublette, 1966: 594
          (syn.) Trissocladius hamiltoni Sæther, 1969: 42
        Hydrobaenus pilipes (Malloch, 1915: 522 as Orthocladius) - Il, OH, ON, widespread
        Hydrobaenus spinnatis Sæther, 1976: 152 - Hudson, pers.comm. 2011

      Hydrosmittia Ferrington & Sæther, 2011: 103

        Hydrosmittia oxoniana (Edwards, 1922: 204 as Camptocladius) - widespread
          (syn.) Parakiefferiella hidakaheia Sasa & Suzuki, 2000b: 189
          (syn.) Prosmittia togarisea (Sasa & Okazawa, 1992)
          (syn.) Pseudosmittia hachijosecunda Sasa, 1994: 47
          (syn.) Pseudosmittia hidakagehea Sasa & Suzuki, 2000b: 188
          (syn.) Pseudosmittia kurobeokasia Sasa & Okazawa, 1992a: 57
          (syn.) Pseudosmittia schachti Casper & Reiss, 1989: 130, pro parte, paratype not holotype
          (syn.) Pseudosmittia togarisea Sasa & Okazawa, 1992b: 160
          (syn.) Pseudosmittia toyamaresea Sasa, 1996a: 39
          (syn.) Pseudosmittia yakyopea Sasa & Suzuki, 2000a: 94
          (syn.) Pseudosmittia yakypequea Sasa & Suzuki, 2000a: 94
          (syn.) Spaniotoma (Smittia) recta Edwards, 1929: 362
        Hydrosmittia ruttneri (Strenzke & Thienemann, 1942: 352 as Pseudosmittia) - Ferrington & Sæther, 2011, Egan & Ferrington 2015: 8 (Isle Royale, pupal exuviae)
          (syn.) Pseudosmittia brevitarsus Brundin, 1947: 40
          (syn.) Pseudosmittia schachti Caspers & Reiss, 1989: 130
          (syn.) Smittia oxoniana Edwards, 1937: 146 not Camptocladius oxonianus Edwards, 1922: 204

      Limnophyes Eaton, 1875: 60

        Limnophyes asquamatus Andersen, 1937: 72 (as L. eltoni var. asquamatus) - BC, WV, widespread Holarctic
          (syn.) Limnophyes septentrionalis Goetghebuer, 1940: 66
          (syn.) Limnophyes smolandicus Brudin, 1947: 32
          (syn.) Limnophyes vernalis Brudin, 1947: 33
          (syn.) Limnophyes franzi Goetghebuer, 1949: 5
          (syn.) Limnophyes hamiltoni Sæther, 1969: 101
        Limnophyes brachytomus (Kieffer, 1922b: 21 as Camptocladius) - MN, widespread
          (syn.) Limnophyes borealis Goetghebuer, 1933: 29
          (syn.) Limnophyes borealis var. squamatus Andersen, 1937: 70
          (syn.) Limnophyes spatulosus Sæther, 1975b: 1055
        Limnophyes carolinensis Sæther, 1975b: 52 - Egan & Ferrington 2015: 9 (Isle Royale, pupal exuviae)
        Limnophyes minimus (Meigen, 1818: 47 as Chironomus) - Sæther 1990 (p. 59; N. Br. Ausable R., Crawford Co.); Egan & Ferrington 2015: 9 (Isle Royale, pupal exuviae)
          (syn.) Limnophyes pusillus Eaton, 1875: 60
          (syn.) Spaniotoma (Limnophyes) pusillus (Eaton), 1875; Edwards, 1929: 355
          (syn.) Limnophyes groenlandiensis Andersen, 1937: 70
          (syn.) Limnophyes interruptus Goetghebuer, 1938: 463
          (syn.) Limnophyes bipunctatus Goetghebuer, 1941: 463
          (syn.) Limnophyes praecox Goetghebuer, 1944: 43
          (syn.) Limnophyes immucronatus Sæther, 1969: 103
          (syn.) Limnophyes hudsoni Sæther, 1975b: 1032
          (syn.) Limnophyes natalensis (Kieffer) sensu Freeman, 1957: 344 (pro parte, misid.)
          (nom. dub.) ?Camptocladius punctulatus Goetghebuer, 1913: 159
          (nom. dub.) ?Camptocladius exiguus Goetghebuer, 1913: 161
          (nom. dub.) ?Camptocladius hexatomus Kieffer, 1915a: 86
          (syn.) ?Limnophyes tamakyioides Sasa, 1983: 79
          (syn.) ?Limnophyes fujinonus Sasa, 1985: 128
        Limnophyes natalensis (Kieffer, 1914: 261 as Camptocladius) - Egan & Ferrington 2015: 9 (Isle Royale, pupal exuviae)
          (syn.) Camptocladius palmensis Santos-Abreu, 1918: 187
          (syn.) Limnophyes brevis Goetghebuer, 1934: 203
          (syn.) Limnophyes bequaerti Goetghebuer, 1939: 61
          (syn.) Limnophyes spinosa Freeman, 1953: 206
          (syn.) Limnophyes nudiradius Sæther, 1975b: 1048
          (nom. dub.) ?Limnophyes jemtlandicus Brundin, 1947: 33
        Limnophyes pumilo (Homgren 1869: 41 as Chironomus) - Egan & Ferrington 2015: 9 (Isle Royale, pupal exuviae)

      Lopescladius Oliveira, 1967: 417

        Lopescladius verruculosus Sæther, 1983b: 289 - Yanoviak & McCafferty 1996 (Mountain Stream, Pine River - Marquette Co.), unassociated larvae and pupae specimens of Lopescladius sp. also taken from Grand Traverse and Newaygo Counties (from sandy substrates)

      Mesosmittia Brundin, 1956: 163

        Mesosmittia patrihortae Sæther, 1985a: 47 - OH
        Mesosmittia prolixa Sæther, 1985a: 48 - IL, OH (expect in s. LP)

      Metriocnemus Wulp, 1874: 136

        Metriocnemus aequalis Johannsen, 1934: 348 - IL, OH
        Metriocnemus albolineatus (Meigen, 1818: 39 as Chironomus) - OH, widespread middle
          (syn.) Metriocnemus atratulus Zetterstedt, 1850: 3590
        Metriocnemus fuscipes (Meigen, 1818: 49 as Chironomus) - widespread north
        Metriocnemus knabi Coquillett, 1904: 11 - E. Bright, collected from pitcher plant Sarracenia purpurea
        Metriocnemus obscuripes (Holmgren, 1869: 38 as Chironomus) - ON, widespread north
          (syn.) Metriocnemus hygropetricus Kieffer, 1912c: 86
          (syn.) Metriocnemus longitarsus Goetghebuer, 1921: 77
        Metriocnemus ursinus (Holmgren, 1869: 39 as Chironomus) - Egan & Ferrington 2015: 9 (Isle Royale, pupal exuviae)

      Nanocladius Kieffer, 1913d: 31

        Nanocladius (Nanocladius) alternantherae Dendy and Sublette, 1959: 510 - OH, widespread southeast
        Nanocladius (Plecopteracoluthus) branchicolus Sæther, 1977: 12- Presque Isle Co. (Rick Jacobsen, pers. comm. 2004)
        Nanocladius (Nanocladius) distinctus (Malloch, 1915: 518 as Orthocladius) - IL, OH, ON widespread
        Nanocladius (Plecopteracoluthus) downesi Steffan, 1965: 1331 - Oliver et al. 1990: 31
        Nanocladius (Nanocladius) incomptus Sæther, 1977b: 43 - OH, ON widespread east
        Nanocladius (Nanocladius) mallochi Sublette, 1970: 70 - OH, IL
          (syn.) Orthocladius distinctus var. bicolor Malloch, 1915: 519, preocc. Zetterstedt, 1838
        Nanocladius (Nanocladius) rectinervis (Kieffer, 1911b: 199 as Cricotopus) - Yanoviak & McCafferty 1996 (Mountain Stream, Huron River - Marquette Co.)
        Nanocladius (Nanocladius) spiniplenus Sæther, 1977: 37 - Egan & Ferrington 2015: 9 (Isle Royale, pupal exuviae)
        Nanocladius (Plecopteracoluthus) sp. 3 - associated with Pteronarcys spp. (Plecoptera: Pteronarcyidae) (Rick Jacobsen, pers. comm. 2004)
        Nanocladius (Plecopteracoluthus) sp. 5 - WI, associated with Nigronia serricornis (Megaloptera: Corydalidae) and Argia spp. (Odonata: Coenagrionidae) (Rick Jacobsen, pers. comm. 2004)

      Orthocladius Wulp, 1874: 132

        Orthocladius (Euorthocladius) abiskoensis Edwards, 1937: 144 - widespread
        Orthocladius (Symposiocladius) annectens Sæther, 1969: 61 - Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
        Orthocladius (Symposiocladius) bilyji Sæther, 2003: 312 - ON, maybe widespread
        Orthocladius (Orthocladius) carlatus (Roback, 1957a: 77 as Hydrobaenus) - Yanoviak & McCafferty 1996 (Salmon-Trout River, Marquette Co.)
        Orthocladius (Orthocladius) clarkei Soponis, 1977: 37 - ON, MN, IL, PA
        Orthocladius (Pogonocladius) consobrinus (Holmgren, 1869: 44 as Chironomus) - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae)
        Orthocladius (Orthocladius) cooki Soponis, 1977: 41 - MN, PA
        Orthocladius (Orthocladius) dentifer Brundin, 1947: 21 - e. ON, widespread west
        Orthocladius (Orthocladius) dorenus (Roback, 1957a: 78 as Hydrobaenus) - Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
          (syn.) Orthocladius currani Sublette, 1967a: 315
        Orthocladius (Eudactylocladius) dubitatus Johannsen, 1942: 72 - Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
        Orthocladius (Orthocladius) frigidus (Zetterstedt, 1838: 811 as Chironomus) - MN (Soponis, 1987)
          (comb.) Spaniotoma (Orthocladius) frigidus Zetterstedt, Edwards, 1929
          (comb.) Hydrobaenus (Bryophaenocladius) frigidus Zetterstedt, Kloet & Hincks, 1945
        Orthocladius (Symposiocladius) lignicola Kieffer in Potthast, 1915: 273 - MN, widespread
        Orthocladius (Euorthocladius) luteipes Goetghebuer, 1938: 457 - widespread Holarctic (expect in w. UP?)
        Orthocladius (Orthocladius) mallochi Kieffer, 1919b: 191 - Soponis, 1977
        Orthocladius (Orthocladius) manitobensis Sæther, 1969: 69 - MN west (expect in w. UP?)
        Orthocladius (Orthocladius) nigritus Malloch, 1915: 191 - Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
        Orthocladius (Orthocladius) obumbratus Johannsen, 1905: 281 - Soponis, 1977, Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
          (syn.) Hydrobaenus paradorenus Roback, 1957a: 79
        Orthocladius (Orthocladius) oliveri Soponis, 1977: 87 - Soponis, 1977
        Orthocladius (Euorthocladius) rivicola Kieffer, 1911b: 181 - Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
        Orthocladius (Euorthocladius) rivulorum Kieffer, 1909: 48 - (coll. EB, Huron River, Washtenaw Co. Feb. 2003)
        Orthocladius (Orthocladius) robacki Soponis, 1977: 92 - Egan & Ferrington 2015: 10 (Isle Royale, pupal exuviae)
        Orthocladius (Euorthocladius) saxosus (Tokunaga, 1939: 326 as Spaniotoma) - widespread Holarctic (expect in w. UP?)
        Orthocladius (Orthocladius) smolandicus Brundin, 1947: 22- MN, widespread north-west (expect in w. UP?)
        Orthocladius (Euorthocladius) thienemanni Kieffer in Kieffer & Thienemann, 1906: 143 - widespread Holarctic

      Parachaetochladius Wülker, 1959: 44

        Parachaetochladius abnobaeus Wülker, 1959 - ON, widespread

      Paracladius Hirvenoja, 1973: 90

        Paracladius alpicola (Zetterstedt, 1850: 3500 as Chironomus) - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae)
        Paracladius quadrinodosus Hirvenoja, 1973: 98) - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae)

      Parakiefferiella Thienemann, 1936a: 135

        Parakiefferiella bathophila (Kieffer, 1912c: 88 as Dactylocladius) - widespread
        Parakiefferiella coronata (Edwards, 1929: 354 as Spaniotoma) - widespread
        Parakiefferiella nigra Brundin, 1949: 827 - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae), widespread Holarctic (Walker et al. 1992)
        Parakiefferiella subaterrima (Malloch, 1915) Camptocladius (orig.) - IL, widespread north
          (syn.) Parakiefferiella torulata Sæther, 1969: 138

      Parametriocnemus Goetghebuer, 1932: 22

        Parametriocnemus lundbeckii (Johannsen, 1905: 302 as Metriocnemus) - Sæther 1969
          (syn.) Metriocnemus innocuus Curran, 1930: 33
          (syn.) Parametriocnemus innocuus (Curran, 1930: 33), Sublette 1966

      Paraphaenocladius Thienemann, 1924: 223

        Paraphaenocladius exagitans (Johannsen, 1905: 303 as Metriocnemus) - IL, OH, WI
          (syn.) Metriocnemus brachyneura Malloch, 1915: 418
          (syn.) Metriocnemus ryutanus Sasa & Hasegawa, 1988: 241
          (syn.) Paraphaenocladius new species 1, Sublette & Sublette 1979: 82
          (syn.) Paraphaenocladius impensus Sasa & Okazawa, 1992a: 56
        Paraphaenocladius impensus (Walker, 1856a: 184 as Chironomus) - MN (expect in UP; widespread northern Holarctic, in seeps, moist soils - as ssp. s. str. - Sæther & Wang 1995)
          (syn.) Chironomus debilipennis Lundbeck, 1898: 286
          (syn.) Metriocnemus angulatus Goetghebuer, 1921: 75
          (syn.) Metriocnemus ampullaceus Kieffer, 1923: 163
          (syn.) Paraphaenocladius albusalatus Chaudhuri & Sinharay, 1987: 95
          (syn.) Paraphaenocladius trichialis Chaudhuri & Sinharay, 1987: 99
          (syn.) Paraphaenocladius distinctus Chaudhuri & Sinharay in Chaudhuri, Bhattacharyay & Dhutta, 1989: 317
          (syn.) Paraphaenocladius amamirobustus Sasa 1990: 131
        Paraphaenocladius innasus Sæther & Wang 1995: 44 - SD-SC, widespread (Sæther and Wang 1995)
        Paraphaenocladius irritus (Walker, 1856: 178 as Chironomus - Sæther & Wang 1995: 20 (as subsp. longiocostatus), probably widespread in North America
        Paraphaenocladius pseudirritus Strenzke, 1950: 216 - Sæther & Wang 1995: 26 (as ssp. nearcticus)
          (syn.) Metriocnemus irratus var. Edwards, 1929: 315
        Paraphaenocladius pusillus Sæther & Wang 1995: 39 - records from South Carolina and Japan may indicate this species has a wide holarctic distribution, or is a species complex

      Parasmittia Strenzke, 1950: 270

        Parasmittia carinata Strenzke, 1950: 270 - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae)

      Psectrocladius Kieffer, 1906: 26

        Psectrocladius (Allopsectrocladius) flavus (Johannsen, 1905: 270 as Orthocladius) - Il, widespread north
        Psectrocladius (Psectrocladius) limbatellus (Holmgren, 1869: 44 as Chironomus) - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae)
          (syn.) Psectrocladius edwardsi Brundin, 1949: 816
        Psectrocladius (Allopsectrocladius) pilosus Roback, 1957a: 88 - Egan & Ferrington 2015: 11 (Isle Royale, pupal exuviae)
        Psectrocladius (Psectrocladius) simulans (Johannsen, 1937a: 67 as Spaniotoma) - Winnell and White 1985
        Psectrocladius (Psectrocladius) subsensilis Sæther and Langton - Egan & Ferrington 2015: 12 (Isle Royale, pupal exuviae)
        Psectrocladius (Psectrocladius) vernalis (Malloch, 1915) Orthocladius (orig.) - IL, MN, OH, ON

      Pseudorthocladius Goetghebuer, 1932: 93

        Pseudorthocladius (Pseudorthocladius) lunatus Sæther & Sublette, 1983: 63 - Oliver et al. 1990:36
        Pseudorthocladius (Pseudorthocladius) uniserratus Sæther & Sublette, 1983: 72 - ON, widespread

      Pseudosmittia Edwards, 1932: 141

        Pseudosmittia digitata Sæther, 1981b: 25 - Ferrington & Sæther, 2011
        Pseudosmittia forcipata (Goetghebuer, 1921: 87 as Camptochironomus) - NB-NM, OH
          (syn.) Smittia triappendiculata Goetghebuer, 1931: 216
          (syn.) Pseudosmittia antillaria Sæther, 1981: 29
          (syn.) Pseudosmittia mongolzeaea Sasa & Suzuki, 1997b: 185
        Pseudosmittia mathildae Albu, 1968: 4 - widespread
          (syn.) Pseudosmittia amamibifurca Sasa, 1990a: 132
          (syn.) Pseudosmittia furudobifurca Sasa & Arakawa, 1994: 100
          (syn.) Pseudosmittia hibaribifurca Sasa, 1993: 80
          (syn.) Pseudosmittia itachibifurca Sasa & Kawai, 1987: 54
          (syn.) Pseudosmittia shoufukuundecima Sasa, 1997: 42
          (syn.) Pseudosmittia trilobata Langton, 1991: 173, not Edwards, 1929: 173
          (syn.) Pseudosmittia sp. B? Strenzke, 1960: 309
          (syn.) Diplosmittia sasai Makarchenko & Makarchenko, 2005: 376
        Pseudosmittia paraconjuncta (Hardy, 1960: 143 as Orthocladius (Smittia) - widespread
          (syn.) Pseudosmittia nr. restricta Sæther, 1977: fig. 60 D-F
        Pseudosmittia ruttneri Strenzke & Thienemann, 1942 - Ferrington & Sæther, 2011

      Psilometriocnemus Sæther, 1969: 105

        Psilometriocnemus triannulatus Sæther, 1969: 107 - MN, ON, widespread

      Rheocricotopus Thienemann & Harnisch, 1932: 135

        Rheocricotopus (Rheocricotopus) effusus (Walker, 1856a: 180 as Chironomus) - IL-SC (Oliver et al. 1990:37)
        Rheocricotopus (Rheocricotopus) eminellobus Sæther, 1969: 85 - ON, widespread (Oliver et al. 1990:37)
        Rheocricotopus (Psilocricotopus) glabricollis (Meigen, 1830: 248 as Chironomus) - MB-NC
        Rheocricotopus (Psilocricotopus) robacki (Beck & Beck, 1964: 204 as Trichocladius) - ON-MT s. to MS-FL, OH (Oliver et al. 1990:37)
          (syn.) Rheocricotopus kenorensis Sæther, 1969: 88

      Rheosmittia Brundin, in Cranston and Sæther 1986: 33

        Rheosmittia unnamed sp. - widespread

      Smittia Holmgren, 1869: 47

        Smittia aterrima (Meigen, 1818: 47 as Chironomus) - UMMZ
        Smittia lasiophtalma (Malloch, 1915: 509 as Camptocladius) - IL
        Smittia lasiops (Malloch, 1915: 508 as Camptocladius) - IL

      Symbiocladius Kieffer, 1925a: 565

        [larvae are parasitic on Ephemeroptera nymphs, usually attached under wing pads of nymph]
        Symbiocladius equitans (Claassen, 1922: 397 as Trissocladius) - widespread (expect in UP)

      Synorthocladius Thienemann, 1935: 211

        Synorthocladius semivirens (Kieffer, 1909: 48 as Dactylocladius) - E. Bright, pers. obs.; Egan & Ferrington 2015: 13 (Isle Royale, pupal exuviae)

      Tavastia Tuiskunen, 1985: 30

        Tavastia cristacauda Sæther, 1991(1992): 386 - White Cedar Swamp, Newago Co. (Sæther, 1991(1992))

      Thienemanniella Kieffer, 1911b: 201

        Thienemanniella elana (Roback, 1957b: 10) Corynoneura (orig.) - OH
        Thienemanniella lobapodema Hestenes & Sæther, 2000: 117 - E. Bright, pers. obs.; Egan & Ferrington 2015: 13 (Isle Royale, pupal exuviae)
          (syn.) Thienemanniella nr. fusca (Kieffer), in Simpson & Bode, 1980
        Thienemanniella norena (Roback, 1957: 38 as Corynoneura) - Egan and Ferrington 2015 (Isle Royale, pupal exuviae)
        Thienemanniella similis (Malloch, 1915: 413) Corynoneura (orig.) - IL, OH, widespread (Oliver et al. 1990:39)
        Thienemanniella taurocapita Hestenes & Sæther, 2000 - E. Bright, pers. obs.
        Thienemanniella xena (Roback, 1957a: 61 as Corynoneura) - E. Bright, pers. obs.

      Tvetenia Kieffer, 1922b: 12

        Tvetenia calvescens (Edwards, 1929: 353 as Spaniotoma) - OH
        Tvetenia paucunca (Sæther, 1969: 46 as Eukiefferiella) - E. Bright, pers. obs.
        Tvetenia tshernovskii (Pankratova, 1968: 253 as Eukiefferiella) - E. Bright, pers. obs.
          (syn.) Eukiefferiella vitracies Sæther, 1969:49
          (syn.) Tvetenia vitracies (Sæther, 1969: 49), Przhibora and Sæther 2010:57, Mason 1985: 418-420
          (syn.) Tvetenia sp. A, Cranston 1982: 73-75
          (syn.) Tvetenia sp. A, Schmid 1993: 208, 491

      Xylotopus Oliver, 1982: 167

        Xylotopus par (Coquillett, 1901: 608 as Orthocladius) - Oliver 1985; Kaufman et al. 1986; Kaufman and King 1987:2280; Michael Kaufman, pers.comm. (February 2013: Augusta Creek, Kalamazoo Co.: 42.37242°N,-85.36052°W; 42.37108°N,-85.36092°W; 42.39061°N,-85.35150°W; 42.39042°N,-85.35135°W)
          (syn.) Eurycnemus scitulus Coquillett, 1901: 608

      Zalutschia Lipina, 1939: 106

        Zalutschia lingulata Sæther, 1976: 222 - Hudson & Bright 2012
        Zalutschia zalutschicola Lipina, 1939: 93 - widespread boreal

    Subfamily PODONOMINAE Thienemann & Edwards, in Thienemann, 1937: 75

      Boreochlus Edwards, 1938: 152 (Tribe Boreochlini)

        Boreochlus persimilis (Johannsen, 1926: 99 as Trichotanypus) - Hudson (2011, pers. comm.)
          (syn.) Boreochlus longisetus Brundin, 1966: 306

      Lasiodiamesa Kieffer, 1924b: 48 (Tribe Boreochlini)

        Lasiodiamesa arientina (Coquillett, 1908: 144 as Tanypus) - ON, WI east (Sæther, 1969:8-15)
          (comb.) Podonomus arientinus (Coquillett), in Johannsen 1934:346, 1952:13
        Lasiodiamesa brusti Sæther, 1969: 15 - MN, widespread northcentral and northwest, may occur in Isle Royale
        Lasiodiamesa tenebrosa (Coquillett, 1905: 66 as Tanypus) - MN, widespread northern
          (syn.) Linacerus piloala Garrett, 1925: 9
          (syn.) Lasiodiamesa nearctica Brundin, 1966: 326
        Lasiodiamesa sphagnicola (Kieffer, 1925c: 16 as Isoplatus) - ON, east

      Paraboreochlus Thienemann, 1939: 166 (Tribe Boreochlini)

        Paraboreochlus stahli Coffman 1988: 192 - KS, Palaearctic

      Parochlus Enderlein, 1912: 109 (Tribe Podonomini)

        Parochlus kiefferi (Garrett, 1925: 8 as Paratanypus) - Egan & Ferrington 2015: 2 (Isle Royale, pupal exuviae)
          (syn.) Tanypus tibialis Staeger, 1845: 354, preocc. Tanypus tibialis Say, 1823)
          (syn.) Paratanypus tibialis Garrett, 1925: 8
          (syn.) Podonomus peregrinus Edwards, 1929: 296
          (syn.) Podonomus sp. Sublette, 1964a: 127

    Subfamily PRODIAMESINAE

      Monodiamesa Kieffer, 1922a: 287

        Monodiamesa bathyphila (Kieffer, 1918a: 102 as Prodiamesa) - Curry 1954: 545 (as P. bathyphila)
          (syn.) Monodiamesa chuzenigra Sasa, 1989
          (syn.) Prodiamesa kamidefea Sasa & Hirabayashi, 1993
        Monodiamesa depectinata Sæther, 1973: 666 - MB, ONT, PA
        Monodiamesa tuberculata Sæther, 1973: 671 - Winnell and White 1985, Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae)

      Odontomesa Pagast, 1947: 502

        Odontomesa fulva (Kieffer, 1919a: 132 as Prodiamesa) - MI - Curry 1954:545 (as Psilodiamesa fulva)
          (syn.) Prodiamesa nigripalpis Goetghebuer, 1922: 52
          (syn.) Monodiamesa flabellata Kieffer, in Zavr̂el, 1926
          (syn.) Cryptochironomus polysetica Botnariuc & Cure, 1956: 259
          (syn.) Orthocladiinae gen. ?1. dospatica Dimitrov, 1963: 13

      Prodiamesa Kieffer, 1906: 37

        Prodiamesa olivacea (Miegen, 1818: 20 as Chironomus) - Curry 1954: 545, Egan & Ferrington 2015: 5 (Isle Royale, pupal exuviae)
          (syn.) Chironomus turpis Zetterstedt, 1838: 811
          (syn.) Tanypus occidentalis Coquillett, 1902b : 92
          (syn.) Chironomus scutellata Meigen, 1818
          (syn.) Diamesa notata Staeger, 1839:
          (syn.) Diamesa consobrinus Zetterstedt, 1850

    Subfamily TANYPODINAE Kieffer, 1906: 33

      Ablabesmyia Johannsen, 1905: 135 (Tribe Pentaneurini Hennig, 1950: 239)

        Ablabesmyia (Asayia) annulata (Say, 1823: 15 as Tanypus) - Roback 1971:378-380
        Ablabesmyia (Ablabesmyia) aspera (Roback, 1959: 124 as Pentaneura) - Roback 1971, Egan & Ferrington 2015: 3 (Isle Royale, pupal exuviae)
          (syn.) Pentaneura gera Roback, 1959
        Ablabesmyia (Ablabesmyia) hauberi Beck & Beck, 1966 - MB-ONT, NY-PA s. to MS-FL
        Ablabesmyia (Ablabesmyia) janta (Roback, 1959: 120) Pentaneura (orig.) - NWT-MD, OH, s. to FL (Roback 1971:372-373)
          (misid.) Pentaneura monilis var. 1, Johannsen 1946:272
        Ablabesmyia (Karelia) illinoensis (Malloch, 1915: 376) Tanypus (orig.) - Roback 1971: 358-360, Egan & Ferrington 2015: 3 (Isle Royale, pupal exuviae)
          (comb.) Pentaneura illinoensis various authors
          (comb.) Pentaneura (Ablabesmyia) illinoensis (Malloch, 1915), Roback 1959:121
          (syn.) Tanypus idei Walley, 1925: 272
        Ablabesmyia (Ablabesmyia) johannseni (Roback, 1959) Pentaneura (orig.) - IL (Roback 1971:371)
          (misid.) Tanypus illinoensis Malloch, 1915 (part):376, paratype 13972
          (syn.) Pentaneura monilis var. 2, Johannsen 1946:273
        Ablabesmyia (Ablabesmyia) mallochi (Walley, 1925: 273) Tanypus (orig.) - Roback 1971:369-371
          (comb.) Pentaneura malloch (Walley, 1925), Hauber 1945: 497, Johannsen 1946: 274
          (misid.) Tanypus illinoensis Malloch, 1915 (part): 376, paratype 13705
          (misid.) Pentaneura illinoensis (Malloch, 1915), Johannsen 1946 (part, var. 6): 273-274
          (syn.) Pentaneura auriensis Roback, 1957a: 39
          (syn.) Pentaneura aequifasciata Dendy & Sublette, 1959: 507
          (syn.) Ablabesmyia ornata Beck & Beck, 1966: 328
          (syn.) Ablabesmyia tarella Roback, 1971: 368
        Ablabesmyia (Ablabesmyia) monilis (Linneaus, 1758: 587) Tipula (orig.) - Roback 1971: 375-378, Egan & Ferrington 2015 (Isle Royale, pupal exuviae)
          (comb.) Ablabesmyia monilis var. nigra Roback, 1971
          (syn.) Tanypus basalis Walley, 1925: 273
          (syn.) nec. Pentaneura basalis, Hauber 1945: 496,498
          (syn.) Pentaneura basalis, Johannsen 1946: 272
          (misid.) nec. Pentaneura basalis?, Roback 1957a: 40, Sublette 1957: 380
          (syn.) Ablabesmyia basalis (Walley, 1925: 273), Roback 1971: 373-374
          (syn.) Ablabesmyia americana Fittkau, 1962: 430 (n. nam. for Tipula monilis, Johannsen, 1905: 142, not Linnaeus 1758, misid.)
        Ablabesmyia (Ablabesmyia) parajanta Roback, 1971 - ONT, NY - NEB s. to FL (Roback 1971:373)
        Ablabesmyia (Karelia) peleensis (Walley, 1926a: 64) Tanypus (orig.) - Roback 1971:361-362
          (misid.) Tanypus illinoensis Malloch, 1915:376, paratype 18799
          (comb.) Pentaneura peleensis (Walley, 1926) various authors
        Ablabesmyia (Karelia) pulchripennis (Lundbeck, 1898: 293 as Tanypus) - QUE-AB,SD-CO
          (syn.) Tanypus prudens Walley, 1925: 275; Walley 1928: 585
          (syn.) Pentaneura prudens (Walley, 1925: 275), Johannsen 1946: 271
          (syn.) Pentaneura (Ablabesmyia) prudens (Walley, 1925: 275), Roback 1959: 120, Dendy and Sublette 1959: 148
          (syn.) Ablabesmyia prudens (Walley, 1925), Sublette and Sublette 1065: 148, Beck and Beck 1966: 320
        Ablabesmyia (Ablabesmyia) rhamphe Sublette 1964:114 - MN-OH s. to FL (Roback 1971:372)
          (syn.) ?Tanypus sp. A, Malloch 1915:397
          (misid.) Pentaneura basalis, Hauber 1945:498 (pupa)
          (misid.) Pentaneura basalis?, Sublette 1957:380, larval ecology

      Apsectrotanypus Fittkau, 1962 (Tribe Macropelopiini Zavrel, 1929: 46)

        Apsectrotanypus johnsoni (Coquillett, 1901: 609 as Tanypus) - Roback 1971

      Brundiniella Roback, 1978: 141 (Tribe Macropelopiini Zavřel, 1929: 46)

        Brundiniella eumorpha (Sublette, 1964a: 107 as Anatopynia) - Roback 1971

      Clinotanypus Kieffer, 1913c: 157 (Tribe Coelotanypodini Fittkau, 1962: 62 Fittkau, 1962: 62)

        Clinotanypus pinguis (Loew, 1861a: 308 as Tanypus) - Roback 1971
          (syn.) Tanypus flavicinctus Loew, 1861: 309
          (syn.) Tanypus thoracicus Loew, 1866: 4
          (syn.) Procladius caliginosus Johannsen, 1905: 131

      Coelotanypus Kieffer, 1913c: 154 (Tribe Coelotanypodini Fittkau, 1962: 62)

        Coelotanypus concinnus (Coquillett, 1895: 308 as Tanypus) - Roback 1971
          (syn.) Coelotanypus flavus Kieffer, 1923a: 296
        Coelotanypus scapularis (Loew, 1866: 2 as Tanypus) - Roback 1971
        Coelotanypus tricolor (Loew, 1861: 309 as Tanypus) -IL, IN, OH

      Conchapelopia Fittkau, 1957: 317 (Tribe Pentaneurini Hennig, 1950: 239)

        Conchapelopia aleta Roback, 1971: 252 - e CA and USA
        Conchapelopia currani (Walley, 1925: 276 as Tanypus) - Cook-MSU
        Conchapelopia fasciata Beck & Beck, 1966: 346 - Egan & Ferrington 2015: 3 (Isle Royale, pupal exuviae)
          (syn.) Conchapelopia dusena Roback, 1971: 247
        Conchapelopia pallens (Coquillett, 1902b: 91 as Tanypus) - AK-QUE, s. to CA-FL
          (syn.) Pentaneura goniodes Sublette, 1964a: 89
          (syn.) Conchapelopia goniodes var. trifida Roback, 1971: 246
        Conchapelopia rurika (Roback, 1957a: 34 as Pentaneura) - MB-QUE, OH, s. to KS-GA
        Conchapelopia telema Roback, 1971: 247 - NWT-ONT, OH, s. to NC

      Djalmabatista Fittkau, 1968: 328 (Tribe Procladiini Roback, 1971: 148)

        Djalmabatista pulchra (Johannsen, 1908: 273 as Protenthes) - Roback 1971:161; Oliver et al. 1990;14
          (syn.) Tanypus apicalis Johannsen, 1908: 64
          (syn.) Protenthes fasciger Curran, 1930: 29
          (syn.) Procladius maculatus Roback, 1971: 152

      Guttipelopia Fittkau, 1962: 251 (Tribe Pentaneurini Hennig, 1950: 239)

        Guttipelopia guttipennis (Wulp, 1861: 16 as Tanypus) - Cook-MSU; see also Bilyj 1988
          (comb.) Tanypus multipunctatus Curran, 1930: 29
          (comb.) Pelpia guttipennis (Wulp), Goetghebuer 1914: 209
          (syn.) Guttipelopia currani Beck & Beck, 1966: 322

      Hayesomyia Murray & Fittkau, 1985: 196 (Tribe Pentaneurini Hennig, 1950: 239)

        Hayesomyia senata (Walley, 1925: 276 as Tanypus) - Cook-MSU
          (comb.) Thienemannimyia (Thienemannimyia) senata (Walley), in Roback 1971
          (syn.) Pentaneura chrysos Sublette, 1964a: 90
          (syn.) ?Chironomus unicolor Walker, 1848: 19 (descr. female)

      Helopelopia Roback, 1971: 250 (Tribe Pentaneurini Hennig, 1950: 239)

        Helopelopia cornuticaudata (Walley, 1925: 277 as Tanypus) - Cook-MSU, Egan & Ferrington 2015: 3 (Isle Royale, pupal exuviae)
        Helopelopia pilicaudata (Walley, 1925: 277 as Tanypus) - NWT-NB, s. to NC

      Labrundinia Fittkau, 1962: 372 (Tribe Pentaneurini Hennig, 1950: 239)

        Labrundinia neopilosella Beck & Beck, 1966: 341 - Cook-MSU
        Labrundinia pilosella (Loew, 1866: 5 as Tanypus) - Cook-MSU; see also Roback, 1987
          (syn.) Labrundinia floridana Beck & Beck, 1966: 339

      Larsia Fittkau, 1962: 339 (Tribe Pentaneurini Hennig, 1950: 239)

        Larsia berneri Beck & Beck, 1966: 350 - MN
        Larsia canadensis Bilyj, 1984: 663 - CAN e. of Rockies
        Larsia decolorata (Malloch, 1915: 370 as Tanypus) - Curry 1954, Roback 1971
          (syn.) Larsia lurida Beck & Beck, 1966: 351
          (syn.) Larsia indistincta Beck & Beck, 1966: 352
        Larsia marginella (Malloch, 1915: 374 as Tanypus) - IL, QUE-KS
        Larsia planensis (Johannsen, 1946: 284 as Pentaneura) - QUE-CA, OH

      Macropelopia Thienemann, 1916: 497 (Tribe Macropelopiini Zavřel, 1929: 46)

        Macropelopia decedens (Walker, 1848: 22 as Tanypus) - Curry 1954
          (syn.) Tanypus hirtipennis Loew, 1866 : 5
          (syn.) Anatopynia dena Roback, in Sublette & Sublette, 1965: 145 [new name for Anatopynia brunnea Roback, 1955: 2]

      Meropelopia Roback, 1971: 253 (Tribe Pentaneurini Hennig, 1950: 239)

        Meropelopia americana (Fittkau, 1957: 320 as Conchapelopia) - Yanoviak & McCafferty 1996 (Pine River, Marquette Co.)
        Meropelopia flavifrons (Johannsen, 1905: 150 as Ablabesmyia) - Roback 1971; Cook-MSU
          (syn.) Pentaneura gesta Roback, in Sublette & Sublette, 1965: 147
          (syn.) Arctopelopia fittkaui Beck & Beck, 1966 : 348
          (comb.) Cantopelopia gesta Roback, in Roback 1971: 254

      Monopelopia Fittkau, 1962: 394 (Tribe Pentaneurini Hennig, 1950: 239)

        Monopelopia boliekae Beck & Beck, 1966: 335, 536 - BC-ONT, s. to MS, FL
        Monopelopia tenuicalcar (Kieffer, 1918a: 181 as Tanypus) - BC-ONT, s. CA-FL
          (syn.) Monopelopia boliekae (in part), Roback, 1971: 280 (Roback, 1986b: 361)

      Natarsia Fittkau, 1962: 151 (Tribe Natarsiini Roback & Moss, 1978)

        Natarsia baltimoreus (Macquart, 1855: 35 as (Coquillett, 1905)Tanypus) - Sublette and Sublette 1965
          (syn.) Ablabesmyia fastuosa Johannsen, 1905: 153

      Nilotanypus Kieffer, 1923b: 191 (Tribe Pentaneurini Hennig, 1950: 239)

        Nilotanypus fimbriatus (Walker, 1848: 20 as Chironomus) - Hudson & Bright 2012
          (syn.) Pentaneura dubia Johannsen, 1946: 286

      Paramerina Fittkau, 1962: 317 (Tribe Pentaneurini Hennig, 1950: 239)

      Pentaneura Philippi, 1865: 629 (Tribe Pentaneurini Hennig, 1950: 239)

        Pentaneura inconspicua (Malloch, 1915: 371 as Tanypus) - IL, ON - widespread
          (syn.) Pentaneura comosa Sublette, 1964a: 93
          (syn.) Pentaneura inculta Beck & Beck, 1966: 334
        Pentaneura inyoensis Sublette, 1964a: 98 - IL, widespread

      Procladius Skuse, 1889: 283 (Tribe Procladiini Roback, 1971: 148)

        Procladius (Holotanypus) abetus Roback, 1971: 185 - Egan & Ferrington 2015: 3 (Isle Royale, pupal exuviae)
        Procladius (Psilotanypus) bellus (Loew, 1866: 4 as Tanypus) - UMMZ; Cook-MSU
          (syn.) Procladius adumbratus Johannsen, 1905: 132
          (syn.) Tanypus pusillus Loew, 1866: 5
          (syn.) Procladius flavidus Kieffer, 1923a: 297
          (syn.) Procladius malifero Garrett, 1925: 10
        Procladius (Holotanypus) culiciformis (Linneaus, 1767: 978 as Tanypus) - Curry 1954 (as Protenthes claripennis), Oliver et al. 1990: 15, Egan & Ferrington 2015 (Isle Royale, pupal exuviae)
          (syn.) Tanypus crassinervis Zetterstedt, 1838: 817
          (syn.) Protenthes claripennis Malloch, 1915: 387
        Procladius (Holotanypus) denticulatus Sublette, 1964a: 124 - Roback 1971
        Procladius (Holotanypus) dentus Roback, 1971: 186 - widespread boreal
        Procladius (Holotanypus) freemani Sublette, 1964: 122 - widespread, mostly boreal
        Procladius (Psilotanypus) macrotrichus Roback, 1971: 168 - MN
        Procladius (Psilotanypus) riparius (Malloch, 1915: 389 as Protenthes) - UMMZ; Cook-MSU
        Procladius (Holotanypus) ruris Roback, 1971: 182 - MN, widespread north
        Procladius (Holotanypus) sublettei Roback, 1971: 191 - Roback 1971

      Psectrotanypus Kieffer, 1909: 42 (Tribe Macropelopiini Zavřel, 1929: 46)

        Psectrotanypus discolor (Coquillett, 1902b) Tanypus (orig.) - WA-NH, WI
        Psectrotanypus dyari (Coquillett, 1902a) Tanypus (orig.) - Roback 1971
          (syn.) Tanypus guttularis Coquillett, 1902b
          (syn.) Tanypus garretti Walley, 1925: 275

      Reomyia Roback, 1986a: 283 (Tribe Pentaneurini Hennig, 1950: 239)

        Reomyia nr. wartinbei (Roback, 1984: 17 as Zavrelimyia) - OH

      Rheopelopia Fittkau, 1962: 209 (Tribe Pentaneurini Hennig, 1950: 239)

        Rheopelopia acra (Roback, 1971) Thienemannimyia (orig.) - Roback 1971
        Rheopelopia paramaculipennis (Roback, 1971) Thienemannimyia (orig.) - OH, IL east
        Rheopelopia perda (Roback, 1971) Thienemannimyia (orig.) - MN, ON widespread north

      Tanypus Meigen, 1803: 261 (Tribe Tanypodini)

        Tanypus (Tanypus) carinatus Sublette, 1964a: 110 - Cook-MSU
        Tanypus (Tanypus) concavus Roback, 1971: 63 - Oliver et al. 1990
        Tanypus (Apelopia) neopunctipennis Sublette, 1964a: 118 - OH, widespread
        Tanypus (Tanypus) punctipennis Meigen, 1818: 61 - Roback 1971
        Tanypus (Tanypus) stellatus Coquillett, 1902b: 89 - Roback 1971
        Tanypus (Tanypus) vilipennis (Kieffer, 1918a: 164 as Protenthes) - Roback 1971; Oliver et al. 1990:16

      Telopelopia Roback, 1971: 255 (Tribe Pentaneurini Hennig, 1950: 239)

        Telopelopia okoboji (Walley, 1928b: 582 as Tanypus) - MB, MN, OH, VA

      Thienemannimyia Fittkau, 1957: 315 (Tribe Pentaneurini Hennig, 1950: 239)

        Thienemannimyia norena (Roback, 1957a: as Pentaneura) - Oliver et al. 1990:14, Egan & Ferrington 2015: 4 (Isle Royale, pupal exuviae)
          (comb.) Thienemannimyia norena var. quadrata Roback, 1971

      Trissopelopia Kieffer, 1923c: 178 (Tribe Pentaneurini Hennig, 1950: 239)

        Trissopelopia ogemawi Roback, 1971 - Oliver et al. 1990:14

      Zavrelimyia Fittkau, 1962: 285 (Tribe Pentaneurini Hennig, 1950: 239)

        Zavrelimyia bifasciata (Coquillett, 1901: 609 as Tanypus) - widespread north and central
        Zavrelimyia sinuosa (Coquillett, 1905 as Tanypus) - Cook-MSU
          (syn.) Zavrelimyia carneosa Fittkau, 1962
          (syn.) Pentaneura fluminalis Sublette, 1964a: 88
        Zavrelimyia thryptica (Sublette, 1964a: 95 as Pentaneura) - ON, widespread

      incerta sedis (Tribe Pentaneurini Hennig, 1950: 239)

        Tanypus futilis Wulp, 1867: 130 - WI

References - Distribution

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Page updates:

  • March 27, 2017 - Reorganized reference sections, splitting them into references pertaining to sources documenting the occurence of a species in Michigan, and another for works that reference authorship of species cited. Corrected omissions and errors in citation.
  • March 22, 2017 - added state records based on Isle Royale NP exuviae collection data in Egan and Ferrington 2015
  • March 17, 2017 - removed Paracladopelma rolli (Chernovskij, 1949), as may be a nomina dubium and its distribution is Palaearctic; its identification in Winnell and White (1985) is problematic
  • June 11, 2016 - Gymnometriocnemus marionensis added to species list based on literature citation (Stur and Ekrem 2015)
  • March 14, 2016 - Paratrichocladius Santos Abreu, 1918, always a difficult genus to differentiate from Cricotopus, has been placed in junoir synonymy with the latter by Cranston and Krosch (2015)
  • March 30, 2015 - Chironomus attenuatus is a nomen dubium (type specimen lost), and (together with its synonomies) was synonomyzed under Chironomus decorus - thanks to Jon Martin for this information
  • December 13, 2013 - added Corynocera sp. based on information from Malcolm Butler and Barbara Hayford (see above)
  • December 11, 2013 - added new state records for Cladopelma galeator, C. amachaerum, and C. collator, removing an erroneous literature citation for C. galeator; it is not in Hudson & Bright 2012
  • February 15, 2013 - removed Chironomus tentans, as it does not occur in North America (except possibly Alaska), instead most specimens identified as that species are Chironomus dilutus Shobanov et al. 1999. Per Martin 2012
  • November 30, 2012 - combined entire Chironomidae list; cleaned up errors in citation, nomenclature
  • November 29, 2012 - removed "working species" from Tanytarsus
  • October 26, 2012 - updated synonomyns for Cladopelma edwardsi and Cladopelma viridulum, added new Michigan records (Hudson & Bright 2012)
  • November 15, 2011 - added Boreochlus persimilis (Johannsen), records from Jordan River, pers. comm. Pat Hudson (2011) - new state record
  • October 24, 2011 - Changed Polypedilum convictum (Walker, 1856) from subgenus Tripodura Townes to Uresipedilum Oyewo & Sæther (Sæther et al. 2010. The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n. Zootaxa 2497: 1–36.)

Page created: December 01, 2002 - Last Updated: April 23, 2017 (EB)