Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Trichoptera (Caddisflies) of Michigan

Introduction

Our current state of knowledge regarding the distribution of Trichoptera in Michigan is improving thorough recent sampling efforts that yield new state records and even new species. Many species still do not have their larval or pupal stages associated with adults, making species-level identifications of immatures difficult or simply not possible.

Previous research by Justin Leonard from the 1930s and 1940s resulted in the first published survey of Trichoptera in Michigan (Leonard and Leonard, 1949). Many subsequent collection and rearing efforts by Justin Leonard principally during the early 1950s, now housed at the University of Michigan Museum of Zoology, Insect Division, and additional collections by others (Pat Hudson, Brian Armitage, Ethan Bright, Doug Bidlack, Ed DeWalt, and Dave Ruiter) added further records to our state's inventory of caddisflies. A new faunal survery of Michigan Trichoptera under the direction of Dr. David Houghton at Hillsdale College has added many further records, and a new publication (Houghton et al. 2016) has since greatly expanded what was known from the Leonards' publication. Based on these efforts and additional literature citations, there are currently about 76 genera and 274 species recorded from Michigan, with another 5 genera and approximately 64 species likely to occur in Michigan based on distributions from nearby localities. Species denoted in bold blue have been recorded in Michigan; other species denoted in bold are likely to occur based on their known distribution. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated. Synonym and literature material information is primarily from the Trichoptera Database Project, a great resource under the directorship of Dr. John Morse at the University of Clemson. You can use the Find function in your web browser to locate families, genera, and species.

    Apataniidae Wallengren, 1884: 128

      Apatania
      Kolenati, 1848: 33
        Apatania incerta (Banks, 1897: 28 as Enoicyla) - PA, WI, MN
        Apatania zonella (Zetterstedt, 1840 as Phryganea stigmatella zonella) - DeWalt & South 2015: 145 (locations 48.09213N -88.59458W and 48.14572N -88.48687W)
          (syn.) Goniotaulus arcticus Boheman, 1865: 568
          (syn.) Apatania groenlandica Kolbe, 1912: 41
          (syn.) Apatania inornata Wallengren, 1886: 78
          (syn.) Apatania palmeni Sahlberg, 1894: 5

    Brachycentridae Ulmer, 1903: 85
    (Photo right: Brachycentrus numerosus larva and case. Copyright 2006 Ethan Bright).

      Brachycentrus Curtis, 1834: 215
        Brachycentrus (Oligoplectrodes) americanus (Banks, 1899: 210 as Oligoplectrum) - Houghton 2016
          (syn.) Oligoplectrodes potanini excisa Martynov, 1928: 473
          (syn.) Oligoplectrodes potanini Martynov, 1910: 356
          (syn.) Brachycentrus similis Banks, 1907a: 124
        Brachycentrus (Sphinctogaster) fuliginosus Walker, 1852: 88
        Brachycentrus (Sphinctogaster) incanus Hagen, 1861: 272
        Brachycentrus (Sphinctogaster) lateralis (Say, 1823: 161 as Phryganea)
        Brachycentrus (Sphinctogaster) numerosus (Say, 1823: 160 as Phryganea)
        Brachycentrus (Sphinctogaster) occidentalis Banks, 1911: 355 - MN, WI (expect near border with UP-MI)

      Micrasema McLachlan, 1876: 253
        Micrasema rusticum (Hagen, 1868: 272 as Dasystoma) - Houghton 2016
          (syn.) Micrasema falcatum Banks, 1914: 265
        Micrasema wataga Ross, 1938a: 178

    Dipseudopsidae Ulmer, 1904: 359

      Phylocentropus
      Banks, 1907a: 130
        Phylocentropus placidus (Banks, 1905: 15 as Holocentropus)
          (syn.) Phylocentropus hansoni Root, 1964: 85
          (syn.) Phylocentropus irroratus Navas, 1934: 20
          (syn.) Phylocentropus maximus Vorhies, 1909: 711
        Phylocentropus lucidus (Hagen, 1861: 294 as Polycentropus) - ONT, PA, OH, IN, IL

    Glossosomatidae Wallengren, 1891: 128

      Agapetus Curtis, 1834: 217
        Agapetus hessi Leonard & Leonard, 1949: 1
        Agapetus tomus Ross, 1941a: 44
        Agapetus walkeri (Betten & Mosely, 1940: 7 as Synagapetus) - ONT, PA, MN
          (syn.) Agapetus rossi Denning, 1941: 200

      Glossosoma Curtis, 1834: 216
        Glossosoma intermedium (Klapálek, 1892: 444 as Mystrophora)
        Glossosoma lividum (Hagen, 1861: 295 as Tinodes)
          (syn.) Glossosoma americanum Banks, 1897: 31
        Glossosoma nigrior Banks, 1911: 355 - Houghton 2016
          (syn.) Eomystra unica Denning, 1942: 46

      Protoptila Banks, 1904a: 215
        Protoptila erotica Ross 1938a: 113 - Houghton 2016
        Protoptila lega Ross, 1941a: 48 - ONT, OH, IL, WI
        Protoptila maculata (Hagen, 1861: 296 as Beraea ?)
          (syn.) Protoptila lloydi Mosely, 1934: 151
          (syn.) Protoptila expositionis Nimmo, 1966: 699
        Protoptila tenebrosa (Walker, 1852: 134 as Hydroptila) - Houghton 2016

    Goeridae Ulmer, 1903: 81

      Goera Stephens, 1829: 28
        Goera calcarata Banks, 1899: 211 - MN, OH Synagapetus (orig.) - ONT, PA, MN
        Goera stylata Ross, 1938a: 172 - Houghton 2016

    Helicopsychidae Ulmer, 1906: 104
    (Photo right: Helicopsyche borealis larva and case. Copyright 2006 Ethan Bright).

      Helicopsyche von Siebold, 1856: 38
        Helicopsyche borealis (Hagen, 1861: 271 as Notidobia) - Houghton 2016
          (syn.) Paludina lustrica Say, 1821: 175 [note: species of Mollusca, suppressed for the purposes of both the Law of Priority and the Law of Homonymy by Opinion 1108 Bulletin of Zoological Nomenclature 35(2): 94-96)]
          (syn.) Valvata arenifera Lea, 1834: 104 [note: [of Mollusca] [doubtful synonym of Helicopsyche borealis (Hagen), Johanson 2002, Insect Systematics and Evolution (Entomologica Scandanavia), Supplement 60: 32, 36-37]
          (syn.) Helicopsyche glabra Hagen, 1864: 130
          (syn.) Helicopsyche californica Banks, 1899: 210
          (syn.) Helicopsyche annulicornis Banks, 1904a: 212

    Hydropsychidae Curtis, 1835

      Arctopsyche McLachlan, 1868: 300 (Arctopsychinae Martynov, 1924b: 25)
        Arctopsyche ladogensis (Kolenati, 1859: 145 as Aphelocheira)
          (syn.) Arctopsyche ladogensis forma obesa McLachlan, 1878: 379
          (syn.) Arctopsyche ramosa form ladogensis Milne, 1936: 67

      Parapsyche Betten, 1934: 181 (Arctopsychinae Martynov, 1924b: 25)
        Parapsyche apicalis (Banks, 1908a: 266 as Arctopsyche)

      Cheumatopsyche Wallengren, 1891: 138 (Hydropsychinae Curtis, 1835: 544)
        Cheumatopsyche analis (Banks, 1903: 243 as Hydropsyche) - Houghton 2016
          (syn.) Cheumatopsyche pettiti Banks, 1908a: 265
        Cheumatopsyche aphanta Ross, 1938a: 151 - new state record, Houghton et al. (2011), Houghton 2016
        Cheumatopsyche campyla Ross, 1938a: 152 - Houghton 2016
        Cheumatopsyche gracilis (Banks, 1899: 216 as Hydropsyche) - Houghton 2016
        Cheumatopsyche lasia Ross, 1938a: 154 - IN, IL, MN
        Cheumatopsyche minuscula (Banks, 1907a: 130) Hydropsyche (orig.)
        Cheumatopsyche oxa Ross, 1938a: 155 - Houghton 2016
        Cheumatopsyche pasella Ross, 1941a: 84 - new state record, Houghton et al. (2011)
        Cheumatopsyche sordida (Hagen, 1861: 290 as Hydropsyche)
        Cheumatopsyche speciosa (Banks, 1904a: 214 as Hydropsyche)
        Cheumatopsyche wabasha Denning, 1947b: 252- OH, WI, MN

      Diplectrona Westwood, 1840 (Diplectroninae Ulmer, 1951: 173)
        Diplectrona modesta Banks, 1908a

      Hydropsyche Pictet, 1834 (Hydropsychinae Curtis, 1835: 544)
        Hydropsyche aerata Ross, 1938a: 144
        Hydropsyche alhedra Ross, 1939a: 67 - Houghton 2016
          (syn.) Hydropsyche racona Denning, 1965: 78
          (syn.) Hydropsyche riola Denning, 1942: 49
          (comb.) Ceratopsyche alhedra (Ross, 1939), various authors
        Hydropsyche alternans (Walker, 1852: 104 as Philopotamus)
          (syn.) Hydropsyche codona Betten, 1934: 187
          (syn.) Philopotamus indecisus Walker, 1852: 104
          (syn.) Philopotamus slossonae var. recurvata Banks, 1914: 253
          (comb.) Ceratopsyche alternans (Walker, 1852), various authors
        Hydropsyche arinale Ross, 1938a: 143 - ONT, IN, IL, WI
        Hydropsyche betteni Ross, 1938a: 146 - Houghton 2016
        Hydropsyche bidens Ross, 1938a: 142
        Hydropsyche bronta Ross, 1938a: 149
          (comb.) Ceratopsyche bronta (Ross, 1938), various authors
        Hydropsyche cheilonis Ross, 1938a: 149
          (comb.) Ceratopsyche cheilonis (Ross, 1938), various authors
        Hydropsyche confusa (Walker, 1852: 103 as Philopotamus) - ONT, OH, MN
          (syn.) Hydropsyche corbeti Nimmo, 1966: 688
          (syn.) Hydropsyche separata Banks, 1936b: 129
        Hydropsyche cuanis Ross, 1938a: 147
        Hydropsyche dicantha Ross, 1938a: 146
        Hydropsyche depravata Hagen, 1861: 290 - IN, OH (southeast, but one record from SK, Canada, hence possibly in Michigan)
        Hydropsyche frisoni Ross, 1938a: 142
        Hydropsyche incommoda Hagen, 1861: 290
        Hydropsyche hageni Banks, 1905: 14 - IL, MN, ON, WI
        Hydropsyche leonardi Ross, 1938a: 145
        Hydropsyche morosa Hagen, 1861: 287
          (syn.) Hydropsyche bifida Banks, 1905: 15
          (syn.) Hydropsyche chlorotica Hagen, 1861: 290
          (comb.) Ceratopsyche morosa (Hagen, 1861), various authors
        Hydropsyche orris Ross, 1938c: 121
          (syn.) Hydropsyche cornuta Ross, 1938a : 141
        Hydropsyche phalerata Hagen, 1861: 287
        Hydropsyche piatrix Ross, 1938a: 148 - widespread: Que-ND-AK
          (comb.) Ceratopsyche piatrix (Ross, 1938), various authors
        Hydropsyche placoda Ross, 1941a: 87 - ONT, OH, IN, IL, WI, MN
        Hydropsyche scalaris Hagen, 1861: 286
        Hydropsyche simulans Ross, 1938a: 139 - IL, IN, OH, ON, MN, WI
        Hydropsyche slossonae Banks, 1908b: 14 - Houghton 2016
          (comb.) Ceratopsyche slossonae (Banks, 1908), various authors
        Hydropsyche sparna Ross, 1938a: 150 - Houghton 2016
          (comb.) Ceratopsyche sparna (Ross, 1938), various authors
        Hydropsyche valanis Ross, 1938a: 144
        Hydropsyche ventura Ross, 1941a: 92 - ONT, PA, MN
          (comb.) Ceratopsyche ventura (Ross, 1941), various authors
        Hydropsyche venularis Banks, 1914: 252 - PA, IL, WI
        Hydropsyche vexa Ross, 1938a: 148 - Houghton 2016
          (comb.) Ceratopsyche vexa (Ross, 1938), various authors
        Hydropsyche walkeri Betten & Mosely, 1940: 23
          (comb.) Ceratopsyche walkeri (Betten & Mosely, 1940), various authors

      Macrostemum Kolenati, 1859: 168 (Macronematinae Ulmer, 1905: 41)
        Macrostemum zebratum (Hagen, 1861: 285 as Macronema)
          (syn.) Macronema australe McLachlan, 1862: 308
          (syn.) Monopseudopsis inscriptus Walker, 1852 : 105

      Potamyia Banks, 1900: 259 (Hydropsychinae Curtis, 1835: 544)
        Potamyia flava (Hagen, 1861) Macronema (orig.)
          (syn.) Hydropsyche kansensis Banks, 1905

    Agraylea larva - copyright 2006 Ethan Bright Hydroptilidae Stephens, 1836: 148
    (Photo right: Agraylea sp. larva, from Rugg Pond, Charlevoix Co., Michigan. Copyright 2006 Ethan Bright).

      Agraylea Curtis, 1834: 217 (Hydroptilinae Stephens, 1836: 148)
        Agraylea costella Ross, 1941b: 15 - ON, WI
        Agraylea multipunctata Curtis, 1834: 217 - Houghton 2016
          (syn.) Hydrochestria argyricola Kolenati, 1948: 35, 104
          (syn.) Allotrichia flavida Banks, 1907b: 164
          (syn.) Agraylea fraterna Banks, 1907b: 164
          (syn.) Agraylea multiguttata Uljanin, 1869: 27, 100
          (syn.) Allotrichia signata Banks, 1904a: 215

      Hydroptila Dalman, 1819: 125 (Hydroptilinae Stephens, 1836)
        Hydroptila ajax Ross, 1938a: 127
        Hydroptila albicornis Hagen, 1861: 275 - IL, IN, MN, OH, WI
        Hydroptila amoena Ross, 1938a: 124 - new state record - Crawford Co. Leonard coll. 1949, Houghton 2016
        Hydroptila ampoda Ross, 1941b: 16 - MN
        Hydroptila angusta Ross, 1938a: 130 - IL, IN, MN, OH, WI
        Hydroptila antennopedia Sykora and Harris, 1994: 68 - MN
        Hydroptila armata Ross, 1938a: 123
        Hydroptila berneri Ross, 1941a: 67
        Hydroptila callia Denning, 1947a: 149
        Hydroptila consimilis Morton, 1905: 65 - Houghton 2016
        Hydroptila delineata Morton, 1905: 66 - IL, IN, MN, OH
        Hydroptila grandiosa Ross, 1938a: 126 - Houghton 2016
        Hydroptila hamata Morton, 1905: 67
        Hydroptila jackmanni Blickle, 1963: 17 - Houghton 2016
        Hydroptila metoeca Blickle & Morse, 1954: 127 - Houghton 2016
        Hydroptila novicola Blickle & Morse, 1954: 124 - Houghton 2016
        Hydroptila perdita Morton, 1905: 67
        Hydroptila quinola Ross, 1947: 147 - Houghton 2016
        Hydroptila salmo Ross, 1941a: 66 - MN, WI
        Hydroptila scolops Ross, 1938a: 128 - new state record - Crawford Co., Leonard coll. 1950
        Hydroptila spatulata Morton, 1905: 66
        Hydroptila strepha Ross, 1941a: 68 - MN, OH, WI
        Hydroptila valhalla Denning, 1947a: 175 - Houghton 2016
        Hydroptila virgata Ross, 1938a: 125 - IL, MN, OH, WI
        Hydroptila waskesia Ross, 1944: 276 - MN, OH
        Hydroptila waubesiana Betten, 1934: 160 - Houghton 2016
        Hydroptila wyomia Denning, 1947a: 150 - Houghton 2016
        Hydroptila xera Ross, 1938a: 132 - new state record, Houghton et al. (2011), Houghton 2016

      Ithytrichia Eaton, 1873: 130 (Hydroptilinae Stephens, 1836: 148; Orthotrichiini Nielsen, 1948: 186)
        Ithytrichia clavata Morton, 1905: 67 - IL, OH, ON, MN, WI (Moulton et al. 1999) [note: larvae of Ithytrichia have been collected from the Crystal River, Leelanau Co., that are probably this species (Craig and Chriscinske 2006]

      Leucotrichia Mosely, 1934: 157 (Leucotrichiinae Flint, 1970: 4)
        Leucotrichia pictipes (Banks, 1911: 359 as Orthotrichia)

      Mayatrichia Mosely, 1937: 182 (Neotrichiinae Ross, 1956)
        Mayatrichia ayama Mosely, 1937: 182

      Neotrichia Morton, 1905: 65 (Neotrichiinae Ross, 1956: 18)
        Neotrichia collata - Morton, 1905: 72
        Neotrichia falca Ross, 1938a: 119 - IL, OH, WI
        Neotrichia halia Denning, 1947a: 153
          (syn.) Neotrichia numii Ross, 1948: 205
        Neotrichia okopa Ross, 1939b: 629 - IL, IN, MN, OH, ON, WI
        Neotrichia vibrans Ross, 1938a: 119 - new state record, Houghton et al. (2011)
          (syn.) Neotrichia ranea Denning, 1947: 20

      Ochrotrichia Mosely, 1934: 162 (Ochrotrichiinae Marshall, 1979: 212)
        Ochrotrichia arva (Ross, 1941a: 58 as Polytrichia)
        Ochrotrichia logana (Ross, 1941a: 54 as Polytrichia)
        Ochrotrichia spinosa (Ross, 1938a: 121 as Polytrichia)
        Ochrotrichia tarsalis (Hagen, 1861: 275 as Hydroptila)
        Ochrotrichia wojcickyi Blickle, 1963: 20 - IN, MN, OH

      Orthotrichia Eaton, 1873: 130 (Orthotrichiinae Nielsen, 1948: 186)
        Orthotrichia aegerfasciella (Chambers, 1873: 114 as Clymene)
          (syn.) Orthotrichia americana Banks, 1904d: 116
          (syn.) Orthotrichia brachiata Morton, 1905: 70
          (syn.) Oxyethira dorsalis Banks, 1904a: 216
        Orthotrichia baldufi Kingsolver & Ross, 1961: 32 - Houghton 2016
        Orthotrichia cristata Morton, 1905: 75 - Houghton 2016
        Orthotrichia curta Kingsolver & Ross, 1961: 33 - Houghton 2016

      Oxyethira Eaton, 1873: 130 (Hydroptilinae Stephens, 1836: 148)
        Oxyethira aeola Ross, 1938a: 117 - new state record, Houghton et al. (2011)
        Oxyethira anabola Blickle, 1966: 185 - MN, ON, WI
        Oxyethira araya Ross, 1941b: 15 - MN, WI
        Oxyethira coercens Morton, 1905: 70 - Houghton 2016
        Oxyethira ecornuta Morton, 1893: 79 - new state record (Alger Co., Ruiter 2011)
        Oxyethira forcipata Mosely, 1934: 153 - Houghton 2016
        Oxyethira grisea Betten, 1934: 162
        Oxyethira michiganensis Mosely, 1934: 153 - Houghton 2016
        Oxyethira novasota Ross, 1944: 135
        Oxyethira obtatus Denning, 1947a: 171 - Houghton 2016
        Oxyethira pallida (Banks, 1904a: 215 as Orthotrichia) - Houghton 2016
        Oxyethira rivicola Blickle & Morse, 1954 - new state record, Houghton et al. (2011), Houghton 2016
          (syn.) Oxyethira cibola Denning, 1948: 12
          (syn.) Oxyethira viminalis Morton, 1905: 71
        Oxyethira serrata Ross, 1938a: 117 - Houghton 2016
        Oxyethira sida Blickle & Morse, 1954: 122 - MN, WI
        Oxyethira verna Ross, 1938a: 118
        Oxyethira zeronia Ross, 1941b: 15 - Houghton 2016

      Stactobiella Martynov, 1924b: 37 (Stactobiinae Botosaneanu, 1956: 382)
        Stactobiella delira (Ross, 1938a: 115 as Stactobia) - new state record, Houghton Co. - Leonard collection
        Stactobiella palmata (Ross, 1938a: 116 as Stactobia) - IL, MN, OH, WI

    Lepidostomatidae Ulmer, 1903: 89

      Lepidostoma Rambur, 1842: 493
        Lepidostoma bryanti (Banks, 1908b: 65 as Alepomyia) - Houghton 2016
          (syn.) Lepidostoma wisconsinense Vorhies, 1909: 685
        Lepidostoma cinereum (Banks, 1899: 210 as Silo)
          (syn.) Lepidostoma strophis Ross, 1938a: 177
        Lepidostoma costale (Banks, 1914: 265 as Olemira)
        Lepidostoma griseum (Banks, 1911: 357 as Phanopsyche)
        Lepidostoma liba Ross, 1941a: 120 - IL, WI
        Lepidostoma prominens (Banks, 1930b: 129 as Arcadopsyche) - WI
        Lepidostoma sackeni (Banks, 1936a: 267 as Mormomyia)
        Lepidostoma sommermanae Ross, 1946: 267 - ONT, NY, OH, IN
        Lepidostoma togatum (Hagen, 1861: 273 as Mormomyia) - Houghton 2016
          (syn.) Pristosilo canadensis Banks, 1899: 212
          (syn.) Silo pallidus Banks, 1897: 29
        Lepidostoma unicolor (Banks, 1911: 357 as Mormomyia)
        Lepidostoma vernale (Banks, 1897: 29 as Mormomyia) - new state record, Houghton et al. (2011)

    Leptoceridae Leach, 1815: 136

      Ceraclea
      Stephens, 1829: 28 (Leptocerinae Leach, 1815: 136; Athripsodini Morse & Wallace, 1976: 34)
        Ceraclea alagma (Ross, 1938a: 155 as Athripsodes) - Houghton 2016
        Ceraclea alces (Ross, 1941a: 95 as Athripsodes) - MN, ON, WI
        Ceraclea ancylus (Vorhies, 1909: 691 as Leptocerus) - IL, IN, MN, OH, ON, WI
        Ceraclea annulicornis (Stephens, 1836: 199 as Leptocerus)
        Ceraclea arielles (Denning, 1942: 48 as Athripsodes)
        Ceraclea cancellata (Betten, 1934: 53 as Leptocerus) - Houghton 2016
        Ceraclea diluta (Hagen, 1861: 277 as Leptocerus)
        Ceraclea erratica (Milne, 1936: 58 as Athripsodes)
        Ceraclea excisa (Morton, 1904: 67 as Leptocerus)
        Ceraclea flava (Banks, 1904a: 212 as Leptocerus)
        Ceraclea maculata (Banks, 1899: 214 as Leptocerus) - Houghton 2016
        Ceraclea mentiea (Walker, 1852: 71 as Leptocerus)
        Ceraclea neffi (Resh, 1974: 269 as Athripsodes)
        Ceraclea nepha (Ross, 1944: 223 as Athripsodes)
        Ceraclea punctuata (Banks, 1894: 180 as Mystacides)
        Ceraclea resurgens (Walker, 1852: 70 as Leptocerus)
        Ceraclea tarsipunctata (Vorhies, 1909: 694 as Leptocerus) - Houghton 2016
        Ceraclea transversa (Hagen, 1861: 279 as Leptocerus) - Houghton 2016
        Ceraclea wetzeli (Ross, 1941a: 94 as Athripsodes) - Houghton 2016

      Leptocerus Leach, 1815: 136 (Leptocerinae Leach, 1815: 136; Leptocerini Leach, 1815: 136)
        Leptocerus americanus (Banks, 1899: 215 as Setodes) - Houghton 2016

      Mystacides Berthold, 1827: 437 (Leptocerinae Leach, 1815: 136; Mystacidini Burmeister, 1839)
        Mystacides interjecta (Banks, 1914: 262 as Oecetina) - Houghton 2016
          (syn.) Mystacides canadensis Banks, 1924: 262
        Mystacides sepulchralis (Walker, 1852: 70 as Leptocerus) - Houghton 2016
          Setodes longula Navas, 1934: 94

      Nectopsyche Mueller, 1879: 40 (Leptocerinae Leach, 1815: 136; Nectopsychini Morse, 1981: 262)
        Nectopsyche albida (Walker, 1852: 71 as Leptocerus) - Houghton 2016
          (syn.) Leptocella coloradensis Banks, 1899: 215
          (syn.) Leptocella gracilis Banks, 1904c: 110
          (syn.) Setodes niveus Hagen, 1861: 281
          (syn.) Mystacides uwarowii Kolenati, 1859: 148
        Nectopsyche candida (Hagen, 1861: 280 as Setodes) - Houghton 2016
        Nectopsyche diarina (Ross, 1944: 218 as Leptocella)
        Nectopsyche exquisita (Walker, 1852: 72 as Leptocerus) - Houghton 2016
          Setodes piffardii McLachlan, 1863: 160
        Nectopsyche pavida (Hagen, 1861: 282 as Setodes) - Houghton 2016

      Oecetis McLachlan, 1877: 294, 329 (Leptocerinae Leach, 1815: 136; Oecetini Silfvenius, 1905: 79)
        Oecetis avara (Banks, 1895: 316 as Setodes)
        Oecetis cinerascens (Hagen, 1861: 282 as Setodes) - Houghton 2016
        Oecetis disjuncta (Banks, 1920: 351 as Oecetina)
        Oecetis immobilis (Hagen, 1861: 283 as Setodes) - Houghton 2016
        Oecetis inconspicua (Walker, 1852: 71-72 as Leptocerus [likely a species complex - see Floyd 1995] - Houghton 2016
        Oecetis nocturna Ross, 1966: 11-14 - IN, MN, OH, ON
        Oecetis ochracea (Curtis, 1825 as Leptocerus.) - MN, OH, ON, WI
          (syn.) Oecetis albescens Mosely, 1930: 240
          (syn.) Oecetis ochracea carri Milne, 1934: 16, 19
          (syn.) Phryganea hectica Zetterstedt, 1840: 137
          (syn.) Mystacida obsoleta Rambur, 1842: 509
          (syn.) Oecetis vicaria Neave, 1934: 162, 169
        Oecetis osteni Milne, 1934: 17, 19 - Houghton 2016
        Oecetis persimilis (Banks, 1907a: 129 as Oecetina) - Houghton 2016

      Setodes Rambur, 1842: 515 (Leptocerinae Leach, 1815: 136; Setodini Morse, 1981: 262)
        Setodes guttatus (Banks, 1900: 257 as Oecetina) - MN, PA
        Setodes incertus (Walker, 1852: 71 as Leptocerus) - Houghton 2016
        Setodes oligius (Ross, 1938a: 160-161 as Leptocerus)

      Triaenodes McLachlan, 1865 (Leptocerinae Leach, 1815: 136; Triaenodini Morse, 1981: 262)
        Triaenodes aba Milne, 1935: 20 - Houghton 2016
        Triaenodes baris Ross, 1938b: 88-89
        Triaenodes dipsius Ross, 1938b: 89 - Houghton 2016
        Triaenodes flavescens Banks, 1900: 257
        Triaenodes frontalis Banks, 1907a: 127 - ON, WI
          (comb.) Ylodes frontalis (Banks, 1907a)
        Triaenodes ignitus (Walker, 1852: 72 as Leptocerus) - Houghton 2016
        Triaenodes injustus (Hagen, 1861: 283 as Setodes) - Houghton 2016
        Triaenodes marginatus Sibley, 1926a: 80 - Houghton 2016
        Triaenodes melacus Ross, 1947: 155-156 - IL, IN, MN, OH, ON, WI
        Triaenodes nox Ross, 1941a: 96 - Houghton 2016
        Triaenodes tardus Milne, 1934: 12, 19 - Houghton 2016
          (syn.) Triaenodes mephitus Milne, 1936: 59
          (syn.) Triaenodes vorhiesi Betten, 1934: 286-287

    Limnephilidae Kolenati, 1848

      Anabolia Stephens, 1837: 204, 229 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848: 30, 38)
        Anabolia bimaculata (Walker, 1852 30 as Limnephilus) - Houghton 2016
        Anabolia consocia (Walker, 1852: 33 as Limniphilus)
          (syn.) Colpotaulius medialis Banks, 1905: 8
          (syn.) Anabolia oslari Ling, 1938: 67
        Anabolia ozburni Milne, 1935: 39, 49
        Anabolia sordida Hagen, 1861: 264

      Asynarchus McLachlan, 1880: 26 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848: 30, 38)
        Asynarchus batchawanus (Denning, 1949: 90-91 as Limnephilus)
          (syn.) Limnephilus conerus Ross, 1950: 426-429
        Asynarchus montanus (Banks, 1907a: 119 as Anabolia) - Houghton 2016
          (syn.) Anabolia curta Banks, 1920: 345
        Asynarchus rossi (Leonard & Leonard, 1949: 4-6 as Limnephilus) [note: species of Asynarchus per D. Ruiter - 1996, contra Wiggins 1996, p. 286]
        Asynarchus mutatus (Hagen, 1861: 267 as Hallesus) - MN, ON

      Frenesia Betten & Mosely, 1940: 165 (Limnephilinae Kolenati, 1848: 30, 35; Chilostigmini Schmid, 1955: 126, 213)
        Frenesia missa (Milne, 1935: 35 as Chilostigma)

      Grammotaulius Kolenati, 1848 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848: 30, 38)
        Grammotaulius interrogationis (Zetterstedt, 1840: 1063 as Phryganea) - MN, ON
          (syn.) Grammotaulius praecox Hagen, 1873a: 451

      Glyphopsyche Banks, 1904b: 141 (Limnephilinae Kolenati, 1848: 30, 35; Chilostigmini Schmid, 1955: 126, 213)
        Glyphopsyche irrorata (Fabricius, 1781: 389 as Phryganea)
          (syn.) Glyphopsyche bryanti Banks, 1904b: 141
          (syn.) Limnephilus intercisus Walker, 1852: 30

      Hesperophylax Banks, 1916: 118 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848)
        Hesperophylax designatus (Walker, 1852: 24 as Limnephilus)
          (syn.) Hesperophylax incisus Banks, 1943b: 348
          (syn.) Hesperophylax minutus Ling, 1938: 67

      Hydatophylax Wallengren, 1891: 31, 73 (Limnephilinae Kolenati, 1848: 30, 35; Stenophylacini Schmid, 1955: 169)
        Hydatophylax argus (Harris, 1869: 333 as Phryganea) - Houghton 2016

      Ironoquia Banks, 1916: 121 (Dicosmoecinae Schmid, 1955: 29; Nothopsychini Vshivkova, Morse, & Ruiter, 2007: 316)
        Ironoquia parvula (Banks, 1900: 256 as Chaetopterygopsis) - DeWalt & South 2015: 147 (location 47.91959N -89.17672W)
        Ironoquia punctatissima (Walker, 1852: 17 as Halesus (orig.)
        Ironoquia lyrata (Ross, 1938a: 163 as Caborius) - Houghton et al. (2011), Houghton 2016

      Lenarchus Martynov, 1914: 222 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848)
        Lenarchus crassus (Banks, 1920: 343 as Limnephilus)

      Leptophylax Banks, 1900: 252 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848)
        Leptophylax gracilis Banks, 1900: 252

      Limnephilus Leach, 1815: 136 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848)
        Limnephilus ademus Ross, 1941b: 18 - new state record (Alger Co., Ruiter 2011)
        Limnephilus argenteus Banks, 1914: 152 - IL, ON, MN, WI
          (syn.) Limnophilus anadyrensis Martynov, 1936: 182
          (syn.) Limnephilus vernalis Nimmo, 1977: 41
        Limnephilus canadensis Banks, 1908a: 164
        Limnephilus dispar McLachlan, 1875: 97
          (syn.) Limnephilus acrocurvus Denning, 1942: 48
          (syn.) Stenophylax minusculus Banks, 1907a: 120
          (syn.) Stenophylax nigridorsus Kolenati, 1848: 23, 32
          (syn.) Limnophilus zhitkovi Martynov, 1914: 197-200
        Limnephilus externus Hagen, 1861: 257 Limnophilus (orig.)
        Limnephilus hyalinus Hagen, 1861: 258 Limnophilus (orig.)
        Limnephilus indivisus Walker, 1852: 34
          (syn.) Limnephilus selatus Denning, 1966: 235
          (syn.) Limnephilus subguttatus Walker, 1852: 34
        Limnephilus infernalis (Banks, 1914: 154 as Anisogamus)
        Limnephilus janus Ross, 1938d: 37 - (Leonard collection, UMMZ)
          (syn.) Colpotaulius minusculus Banks, 1924: 439
        Limnephilus moestus Banks, 1908b: 61 - Houghton 2016
        Limnephilus ornatus Banks, 1897: 27 - Houghton 2016
        Limnephilus partitus Walker, 1852: 32 - MN, ON
        Limnephilus parvulus (Banks, 1905: 9 as Stenophylax?)
        Limnephilus perpusillus Walker, 1852: 35
        Limnephilus rhombicus (Linnaeus, 1758: 548 as Phryganea) - Houghton 2016
          (syn.) Limnephilus chilcotinensis Nimmo, 1991: 33
          (syn.) Limnephilus combinatus Walker, 1852: 28
          (syn.) Phryganea rhomboidica Berkenhout, 1795: 294
        Limnephilus sackeni Banks, 1930a: 227
        Limnephilus samoedus (McLachlan, 1880: 53 as Astratus)
          (syn.) Zaporota pallens Banks, 1920: 342
          (syn.) Limnephilus pallens (Banks, 1920: 342)
        Limnephilus secludens Banks, 1914: 152
        Limnephilus sericeus (Say, 1824: 309 as Phryganea) - Houghton 2016
          (syn.) Anabolia decepta Banks, 1899: 208
          (syn.) Limnephilus despectus Walker, 1852: 31
          (syn.) Limnephilus eminens Betten, 1934: 323
          (syn.) Apatania fuscostigma Matsumura, 1931: 1123
          (syn.) Limnephilus multifarius Walker, 1852: 32
          (syn.) Limnephilus perforatus Walker, 1852: 33
          (syn.) Phryganea pilosula Zetterstedt, 1840: 90
          (syn.) Limnophilus shimushirensis Tsuda, 1942: 316
        Limnephilus sublunatus Provancher, 1877: 243 as Limnophilus (spell.) - MN, ON
          (syn.) Limnophilus americanus Banks, 1900: 253
          (syn.) Limnephilus macgillivrayi Banks, 1908a: 263
        Limnephilus submonilifer Walker, 1852: 33

      Nemotaulius Banks, 1906: 107 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848)
        Nemotaulius hostilis (Hagen, 1873a: 444 as Glyphotaelius) - Houghton 2016

      Onocosmoecus Banks, 1943b: 357 (Dicosmoecinae Schmid, 1955: 29)
        Onocosmoecus unicolor (Banks, 1897: 27 as Anabolia)
          (syn.) Dicosmoecus alascensis Banks, 1943b: 365
          (syn.) Dicosmoecus coloradensis Ulmer, 1905a: 64
          (syn.) Dicosmoecus flavus Martynov, 1914: 253
          (syn.) Dicosmoecus occidentis Banks, 1943b: 362
          (syn.) Anabolia quadrinotata Banks, 1908b: 62 (as Anabolia 4-notata)
          (syn.) Asynarchus tristis Banks, 1900: 254

      Phanocelia Banks, 1943b: 354 (Limnephilinae Kolenati, 1848: 30, 35; Chilostigmini Schmid, 1955: 126, 213)
        Phanocelia canadensis (Banks, 1924: 442 as Apatania) - specimens from ESGR in 1950s, in UMMZ-Insect Division.

      Platycentropus Ulmer, 1905b: 13 (Limnephilinae Kolenati, 1848: 30, 35; Limnephilini Kolenati, 1848)
        Platycentropus amicus (Hagen, 1861: 265 as Hallesus)
          (syn.) Platycentropus plectrus Ross, 1938a: 169
        Platycentropus indistinctus (Walker, 1852: 37 as Limnephilus)
          (syn.) Hylepsyche fraternus Banks, 1943b: 349
        Platycentropus radiatus (Say, 1824: 308 as Phryganea) - Houghton 2016
          (syn.) Halesus hostis Hagen, 1861: 266
          (syn.) Limnephilus indicans Walker, 1852: 23
          (syn.) Hallesus maculipennis Kolenati, 1859: 159, 176, 280

      Pseudostenophylax Martynov, 1909b (Pseudostenophylacinae Schmid, 1955: 19)
        Pseudostenophylax sparsus (Banks, 1908a: 63 as Halesus) - Houghton 2016
          (syn.) Stenophylax calypso Banks, 1911: 350

      Psychoglypha Ross, 1944: 180, 201 (Limnephilinae Kolenati, 1848: 30, 35; Chilostigmini Schmid, 1955: 126, 213)
        Psychoglypha subborealis (Banks, 1924: 441 as Chilostigma)

      Pycnopsyche Banks, 1905: 9 (Limnephilinae Kolenati, 1848: 30, 35; Stenophylacini Schmid, 1955: 19)
        Pycnopsyche aglona Ross, 1941b: 18
        Pycnopsyche antica (Walker, 1852: 9 as Neuronia) - Houghton 2016
          (syn., in part.) Pycnopsyche minima Banks, 1943b: 345
        Pycnopsyche circularis (Provancher, 1877: 259 as Platyphylax) - OH, WI
        Pycnopsyche guttifer (Walker, 1852: 16 as Halesus) - Houghton 2016
          (syn.) Pycnopsyche similis Banks, 1907a: 122
        Pycnopsyche indiana (Ross, 1938c: 121 as Stenophylax) - Houghton et al. 2009
        Pycnopsyche lepida (Hagen, 1861: 269 as Enoicyla) - Houghton 2016
        Pycnopsyche limbata (McLachlan, 1871: 108 as Stenophylax) - MN, ON, WI
        Pycnopsyche scabripennis (Rambur, 1842: 488 as Limnephila)
          (syn., in part) Pycnopsyche minima Banks, 1943b: 345
          (syn.) Pycnopsyche perplexa Betten & Mosely, 1940: 149
        Pycnopsyche subfasciata (Say, 1828 as Phryganea) - Houghton 2016

    Molannidae Wallengren, 1891: 116

      Molanna
      Curtis, 1834: 214
        Molanna blenda Sibley, 1926b: 210 - Houghton 2016
        Molanna flavicornis Banks, 1914: 261
        Molanna tryphena Betten, 1934: 245 - Houghton 2016
        Molanna ulmerina Navas, 1934: 23
          (syn.) Molanna musetta Betten, 1934: 245
        Molanna uniophila Vorhies, 1909: 705 - Houghton 2016, Mundahl & Mundahl 2015 (44.0245°N, -84.5482°W, Pratt Lake, Gladwin Co.)

    Odontoceridae Wallengren, 1891: 12

      Marilia
      Mueller, 1880: 127 (Odontocerinae Wallengren, 1891: 12)
        Marilia flexuosa Ulmer, 1905a: 70 - IN, ON
          (syn.) Anisocentropus fuscus Banks, 1905: 19

      Psilotreta
      Banks, 1899: 213 (Odontocerinae Wallengren, 1891: 12)
        Psilotreta indecisa (Walker, 1852: 95 as Goera)
          (syn.) Heteroplectron borealis Provancher, 1877: 263

    Philopotamidae Stephens, 1829

      Chimarra Stephens, 1829: 318 (Chimarrinae Rambur, 1842: 498)
        Chimarra aterrima Hagen, 1861: 297 as Chimmarrha (spell.)
          (syn.) Wormaldia buenoi Navas, 1934: 82
        Chimarra feria Ross, 1941a: 51 as Chimarrha (spell.) - Houghton 2016
        Chimarra obscura (Walker, 1852: 121 as Beraea?) - Houghton 2016
          (syn.) Chimarrha lucia Betten, 1934: 175
          (syn.) Wormaldia plutonis Banks, 1911: 358
        Chimarra socia Hagen, 1861: 297 as Chimarrha (spell.)
          (syn.) Wormaldia femoralis Banks, 1911: 358

      Dolophilodes Ulmer, 1909: 125 (Philopotaminae Stephens, 1829: 316)
        Dolophilodes distincta (Walker, 1852: 104 as Philopotamus) - Houghton 2016
          (comb.) Trentonius distinctus Betten & Mosely, 1940
          (syn.) Philopotamus americanus Banks, 1895: 316

      Wormaldia McLachlan, 1865: 140 (Philopotaminae Stephens, 1829: 316)
        Wormaldia moesta (Banks, 1914: 202 as Paragapetus)
          (syn.) Dolophilus breviatus Banks, 1914: 254

    Phryganeidae Leach, 1815 (all Michigan genera in Subfamily Phryganeinae Leach, 1815)

      Agrypnia Curtis, 1835
        Agrypnia colorata (Hagen, 1873a) Agrypnia (orig.)
          (syn.) Prophryganea bradorata Milne, 1931
          (syn.) Phryganea principalis Martynov, 1909a
        Agrypnia deflata (Milne, 1931) Prophryganea (orig.) - ON
          (syn.) Dasystegia (Dasystegia) obsoleta deflata Milne, 1934
        Agrypnia glacialis Hagen, 1873b (orig.) - MN, ON, WI
          (syn.) Asynarchus alascensis Banks, 1900
          (syn.) Agrypnia dextra Ross, 1938a
        Agrypnia improba (Hagen, 1873a) Phryganea (orig.)
          (syn.) Dasystegia improba sackeni Banks, 1943b
        Agrypnia macdunnoughi (Milne, 1931 ) Prophryganea (orig.) - (Wiggins 1998), Bass L., Marquette Co., 22. vi. 1928 (P); Copper Harbor, Keweenaw Co. 26.vi.1957 - records at MSU (R. Fischer)
        Agrypnia straminea Hagen, 1873a
          (syn.) Agrypnetes curvata Banks, 1900
          (syn.) Phryganomyia obscura Banks, 1907a
        Agrypnia vestita (Walker, 1852) Neuronia (orig.)
          (syn.) Neuronia commixta Walker, 1852

      Banksiola Martynov, 1924a
        Banksiola crotchi Banks, 1943a - Houghton 2016
          (syn.) Banksiola selina Betten, 1944
        Banksiola smithi (Banks, 1914) Neuronia (orig.)
        Banksiola dossuaria (Say, 1828) Phryganea (orig.) - Houghton et al. (2011), Houghton 2016

      Beothukus Wiggins, 1989
        Beothukus complicatus (Banks, 1924) Ecclisomyia (orig.)

      Fabria Milne, 1934
        Fabria inornata (Banks, 1907a) Neuronia (orig.)

      Hagenella Martynov, 1924a
        Hagenella canadensis (Banks, 1907a) Neuronia (orig.) - Houghton 2016

      Oligostomis Kolenati, 1848
        Oligostomis ocelligera (Walker, 1852) Neuronia (orig.)
          (syn.) Neuronia stygipes Hagen, 1873a
        Oligostomis pardalis (Walker, 1852) Neuronia (orig.) - OH, ON

      Phryganea Linnaeus, 1758
        Phryganea cinerea Walker, 1852 - Houghton 2016
          (syn.) Phryganea divulsa Walker, 1860
        Phryganea sayi Milne, 1931
          (syn.) Phryganea interrupta Say, 1828

      Ptilostomis Kolenati, 1859
        Ptilostomis angustipennis Hagen, 1873a
        Ptilostomis ocellifera (Walker, 1852) Neuronia (orig.) - Houghton 2016
          (syn.) Ptilostomis kovalevskii variety b Kolenati, 1859
          (syn.) Neuronia simulans Betten & Mosely, 1940
        Ptilostomis postica (Walker, 1852) Neuronia (orig.)
          (syn.) Ptilostomis kovalevskii variety a Kolenati, 1859
        Ptilostomis semifasciata (Say, 1828) Phryganea (orig.) - Houghton 2016
          (syn.) Neuronia dubitans Betten & Mosely, 1940
          (syn.) Neuronia fusca Walker, 1852

    Polycentropodidae Ulmer, 1903: 11, 31, 33 117
    (All Michigan genera of Polycentropodidae are grouped in the subfamily Polycentropodinae Ulmer, 1903)

      Cernotina Ross, 1938a: 136
        Cernotina spicata Ross, 1938a: 138 - Houghton 2016

      Cyrnellus Banks, 1913: 88
        Cyrnellus fraternus (Banks, 1905: 17 as Cyrnus)

      Holocentropus McLachlan, 1878: 375, 400
        Holocentropus flavus Banks, 1908b: 66 - Houghton 2016
        Holocentropus glacialis Ross, 1938a: 135
        Holocentropus interruptus Banks, 1914: 257
          (syn.) Holocentropus orotus Banks, 1914: 257
          (syn.) Holocentropus longus Banks, 1914: 257
        Holocentropus melanae Ross, 1938a: 136

      Neureclipsis McLachlan, 1864: 26, 30
        Neureclipsis bimaculata (Linnaeus, 1758: 548 as Phryganea) - Houghton 2016
          (syn.) Polycentropus concolor Burmeister, 1839: 914
          (syn.) Phryganea noctuaeformis von Paula Schrank, 1802: 180
          (syn.) Anticyra? robusta Walker, 1852: 122
          (syn.) Phryganea tigurinensis Fabricius, 1798: 34
          (syn.) Philopotamus variegatus Schoch, 1884: 50-52
        Neureclipsis crepuscularis (Walker, 1852: 87 as Brachycentrus) - Houghton 2016
        Neureclipsis valida (Walker, 1852: 100 as Polycentropus)

      Nyctiophylax Brauer, 1865: 419
        Nyctiophylax affinis (Banks, 1897: 30 as Polycentropus) - new state record, Houghton et al. (2011), Houghton 2016
          (syn.) Nyctiophylax zelenus Denning, 1950: 99
          (nom. dub.) Nyctiophylax vestitus Hagen, 1861: 293
          (comb.) Polycentropus vestitus Hagen, 1860
        Nyctiophylax banksi Morse, 1972: 173 - ONT, PA, MN
        Nyctiophylax celta Denning, 1947b: 251 - ONT, PA, IL, WI, MN
        Nyctiophylax moestus Banks, 1911: 359 - Houghton 2016
        Nyctiophylax uncus Ross, 1944: 70

      Phylocentropus Banks, 1907a: 130 (see family Dipseudopsidae)

      Plectrocnemia Stephens, 1836: 167
        [Note: Plectrocnemis reinstated as a full genus by Commorra & Holzenthal 2011)
        Plectrocnemia albipuncta Banks, 1930b: 131 - Houghton 2016
          (comb.) Polycentropus albipunctus (Banks, 1830)
        Plectrocnemia aureola Banks, 1930b: 130 - Houghton 2016
          (comb.) Polycentropus aureolus (Banks, 1930b)
        Plectrocnemia cinerea (Hagen, 1861: 293 as Polycentropus) - Houghton 2016
          (syn.) Polycentropus canadensis Banks, 1897: 31
          (syn.) Holocentropus flavicornis Banks, 1907b: 162
          (syn.) Plectrocnemia lutea Betten, 1934: 219
          (syn.) Plectrocnemia pallescens Banks, 1930a: 231
        Plectrocnemia clinei Milne, 1936: 87- Houghton 2016
          (comb.) Polycentropus clinei (Milne, 1936)
        Plectrocnemia crassicornis (Walker, 1852: 101 as Polycentropus)
          (syn.) Plectrocnemia adironica Banks, 1914: 256
          (syn.) Polycentropus confusus Hagen, 1861: 293
          (syn.) Plectrocnemia australis Banks, 1907a : 131
        Plectrocnemia nascotius Ross, 1941a: 73 as Polycentropus) - new state record (Alger Co., Ruiter 2011)
        Plectrocnemia remota (Banks, 1911: 359 as Polycentropus) - Houghton 2016
        Plectrocnemia sabulosus (Leonard & Leonard, 1949: 3 as Polycentropus)
        Plectrocnemia weedi (Blickle & Morse, 1955: 95 as Polycentropus) - Houghton 2016

      Polycentropus Curtis, 1835: 544
        [Note: Plectrocnemia reinstated as genus by Commorra & Holzenthal 2011, 9 species previously in Polycentropus moved]
        Polycentropus centralis Banks, 1914: 258
        Polycentropus confusus Hagen, 1861: 293
        Polycentropus elarus Ross, 1944: 59, 64-66 - ONT, OH, IN
        Polycentropus milaca Etnier, 1968: 189 - MN
        Polycentropus pentus Ross, 1941a: 71 - Houghton 2016
        Polycentropus picicornis Stephens, 1836: 177 as Neureclipsis) - MN
          (syn.) Polycentropus irroratus Stephens, 1836: 178
          (syn.) Plectrocnemia liturata Kolenati, 1859: 145
          (syn.) Cyrnus pulchellus Stephens, 1836: 175
          (syn.) Polycentropus subpunctatus Stephens, 1836: 176
        Polycentropus timesis (Denning, 1948a: 119 as Neureclipsis) - new state record, Houghton et al. (2011)
          (in parte) Polycentropus picicornis Stephens, 1836

    Psychomyiidae Walker, 1852: 80

      Lype McLachlan, 1878: 409, 422 (Tinodinae Li & Morse, 1997)
        Lype diversa (Banks, 1914: 253 as Psychomyia) - Houghton 2016
          (syn.) Lype griselda Betten, 1934: 229

      Psychomyia Latreille, 1829 (Psychomyiinae Walker, 1852)
        Psychomyia flavida Hagen, 1861: 294 - Houghton 2016
          (syn.) Psychomyiella composita Martynov, 1910: 408-411
          (syn.) Psychomyia moesta Banks, 1907a: 131-132
          (syn.) Psychomyia pulchella Banks, 1899: 217

    Rhyacophilidae Stephens, 1836: 148

      Rhyacophila Pictet, 1834: 23, 181
        Rhyacophila brunnea Banks, 1911 ("1910"): 352-353, 355
          (syn.) Rhyacophila acuminata Fields, 1981: 502
          (syn.) Rhyacophila acropedes Banks, 1914: 302
          (syn.) Rhyacophila ignorata Schmid, 1974: 933
        Rhyacophila fenestra Ross, 1938a: 102 - OH, IN, IL
        Rhyacophila fuscula (Walker, 1852: 10 as Neuronia)
        Rhyacophila glaberrima Ulmer, 1907: 85 - OH, IN, IL
          (syn.) Rhyacophila andrea Betten, 1934: 127
          (syn.) Rhyacophila fairchildi Banks, 1930: 130
        Rhyacophila invaria (Walker, 1852: 101 as Polycentropus) - ONT, WI
          (syn.) Rhyacophila luctuosa Banks, 1911 ("1910"): 351
        Rhyacophila ledra Ross, 1939a: 65
          (syn.) Rhyacophila hardeni Denning, 1948b: 101
        Rhyacophila lobifera Betten, 1934: 131 - Bruce McCulloch 2013, pers.comm.
        Rhyacophila mainensis Banks, 1911 ("1910"): 354 - Houghton 2016
          (syn.) Rhyacophila melita Ross, 1938a: 104
        Rhyacophila manistee Ross, 1938a: 104 - Houghton 2016
        Rhyacophila torva Hagen, 1861: 296 - PA, OH, WI
          (syn.) Rhyacophila terminata Banks, 1907a: 132
          (syn.) Rhyacophila vinura Milne, 1936: 100
        Rhyacophila vibox Milne, 1936: 104 - Houghton 2016

    Sericostomatidae Stephens, 1836: 148, 180
      Agarodes Banks, 1899: 217
        Agarodes distinctus (Ulmer, 1905a: 67-68 as Notidobia)
          (syn.) Schizopelex lobata Banks, 1911: 356
          (syn.) Psiloneura moesta Banks, 1914: 264

    Uenoidae Iwata, 1927: 214

      Neophylax McLachlan, 1871 (Thremmatinae Martynov, 1935: 386)
        Neophylax aniqua Ross, 1947: 153 - ON (expect in the n. and e. UP)
        Neophylax ayanus Ross, 1938a: 168
        Neophylax concinnus McLachlan, 1871: 111 - Houghton 2016
          (syn.) Neophylax autumnus Vorhies, 1909: 669-672
        Neophylax fuscus Banks, 1903: 242
        Neophylax oligius Ross, 1938a: 168 - Houghton 2016

      Oligophlebodes Ulmer, 1905a: 66 (Thremmatinae Martynov, 1935: 386)
        Oligophlebodes minutus (Banks, 1897: 28 as Halesus)
          (syn.) Oligophlebodes coloradensis Ulmer, 1905a: 66
          (syn.) Apatania pictula Banks, 1943b: 355

References - Distribution

    Chamorro ML, Holzenthal RW. 2011.
    Phylogeny of Polycentropodidae Ulmer, 1903 (Trichoptera: Annulipalpia: Psychomyioidea) inferred from larval, pupal and adult characters. Invertebrate Systematics 25: 219-253. PDF
    Craig JM, Chriscinski MA. 2006. First record of Ithytrichia (Trichoptera: Hydroptilidae) in Michigan. The Great Lakes Entomologist 118: 313-314.
    Etnier DA, Parker CR, Baxter JT, Long TM. 2010. A review of the genus Agapetus Curtis (Trichoptera: Glossosomatidae) in eastern and central North America, with description of 12 new species. Insecta Mundi 0149:1-77. PDF
    DeWalt RE, South EJ. 2015. Ephemeroptera, Plecoptera, and Trichoptera on Isle Royale National Park, USA, compared to mainland species pool and size distribution. ZooKeys 532: 137-158.
    Flint OS. 1984. The genus Brachycentrus in North America, with a proposed phylogeny of the genera of Brachycentridae (Trichoptera). Smithsonian Contributions to Zoology 398. 58 pp. PDF
    Gordon AE. 1974. A synopsis and phylogenetic outline of the nearctic members of Cheumatopsyche. Proceedings of the Academy of Natural Sciences of Philadelphia 126(9):117-160.
    Houghton DC, Brandin CM, Brakel KA. 2011. Analysis of the caddisflies (Trichoptera) of the Manistee River watershed, Michigan. The Great Lakes Entomologist 44(1): 1-15. PDF
    Houghton DC, Lux C, Spahr M. 2009. Demonstration of sex pheromones in Molanna uniophila (Trichoptera: Molannidae), Platycentropus radiatus, Pycnopsyche indiana, and P. subfasciata (Trichoptera: Limnephilidae), with an assessment of interspecific attraction between four sympatric Pycnopsyche species. The Great Lakes Entomologist 42(1-2):72-77. PDF.
    Houghton DC. 2016. The caddisflies (Trichoptera) of an undisturbed lower Michigan habitat. The Great Lakes Entomologist 49(1-2): 41-54. PDF.
    Houghton DC, Pytel AJ, Dewalt RE, Brandin CM, Rogers SE, Armitage BJ, Hudson PL, Ruiter DE, Bright E. 2016. Updated checklist of the Michigan (USA) caddisflies, with regional and habitat affinities. Proceedings of the XV International Symposium on Trichoptera. submitted, pending publication.
    Leonard JW, Leonard FA. 1949. Noteworthy records of caddis flies from Michigan, with descriptions of new species. Occassional Papers of the Museum of Zoology, University of Michigan 520. 20p. PDF.
    Moulton SR, Harris SJ, Slusark JP. 1999. The microcaddisfly genus Ithytrichia Eaton (Trichoptera: Hydroptilidae) in North America. Proceedings of the Entomological Society of Washington 101(2): 233-241. PDF.
    Mundahl ND, Mundahl ED. 2015. Habitats and foraging movements of larvae of Molanna uniophila Vorheis (Trichoptera: Molannidae) in Pratt Lake, Michigan, USA. Journal of Insects 2015: 1-8.
    Ross HH. 1938a. Descriptions of Nearctic caddis flies (Trichoptera) with special reference to the Illinois species. Bulletin of the Illinois Natural History Survey 21: 101-183. PDF.
    Ross HH. 1944. The caddisflies, or Trichoptera, of Illinois. Bulletin of the Illinois Natural History Survey 23: 1-326. PDF
    Ruiter, DE. 2000. New descriptions and distributional records for Eastern North American caddisflies (Trichoptera). Entomological News 111(3):227-232.
    Vineyard RN, Wiggins GB, Frania HE, Schefter PW. 2005. The caddisfly genus Neophylax (Trichoptera: Uenoidae) . Royal Ontario Museum Contributions in Science 2. v + 141 p.
    Wiggins GB. 1996. Larvae of the North American caddisfly genera (Trichoptera), 2nd Edition. University of Toronto Press: Toronto, Canada. Wymer DA, Morse JC. 2000. Larva, pupa, and adults of Glossosoma nigrior (Trichoptera: Glossosomatidae) with a review of the eastern North American species of Glossosoma. Entomological News 111(3):149-158.
References - Taxonomic

    Banks N. 1894. On a collection of neuropteroid insects from Kansas. Entomological News 5: 178-180. PDF.
    Banks N. 1895. New neuropteroid insects. Transactions of the American Entomological Society 22: 313-316. PDF.
    Banks N. 1897. New North American neuropteroid insects. Transactions of the American Entomological Society 24: 21-31. PDF.
    Banks N. 1899. Descriptions of new North American neuropteroid insects. Transactions of the American Entomological Society 25: 199-218. PDF.
    Banks N. 1900. New genera and species of Nearctic neuropteroid insects. Transactions of the American Entomological Society 26: 239-260. PDF.
    Banks N. 1903. Some new neuropteroid insects. Journal of the New York Entomological Society 11: 236-243. PDF.
    Banks N. 1904a. A list of neuropteroid insects, exclusive of Odonata, from the vicinity of Washington, D.C. Proceedings of the Entomological Society of Washington 6: 201-217.
    Banks N. 1904b. Two new species of caddice-flies. Proceedings of the Entomological Society of Washington 6: 140-142.
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Page edits:
    March 27, 2017
    • Reorganized reference sections, splitting them into references pertaining to sources documenting the occurence of a species in Michigan, and another for works that reference authorship of species cited. Corrected omissions and errors in citation and systematic placement.
    March 24, 2017
    • Incorporated systematic changes in Geraci et al. 2010 that places Ceratopsyche Ross & Unzicker, 1977 as a junior synonym of Hydropsyche Pictet, 1834
    March 12, 2017
    • Incorporated further systematic changes in Polycentropodidae as noted in Chamorro & Holzenthal (2011). Holocentropus resurrected as a valid genus, as Polycentropus, as previously understood, was not monophyletic
    February 23, 2015
    • DeWalt & South 2015 records Apatania zonella and Ironoquia parvula from Isle Royale
    November 17, 2013
    • Bruce McCulloch records Rhyacophila lobifera from the Rouge River watershed in southeastern Michigan
    November 06, 2012
    • Chamorro & Holzenthal (2011) has species formerly placed in Polycentropus - albipunctus Banks, aureolus (Banks), cinereus (Hagen), clinei Milne, crassicornis Walker, 1852, nascotius Ross, remotus Banks, sabulosus Leonard & Leonard, 1949, and weedi Blickle & Morse, 1955 - moved (in some cases reinstated) to Plectrocnemia.
    November 04, 2012
    • Beginning to keep track of changes to the web site pages
    • Removed individual pages for families into one long list, with greatly streamlined html coding. Families accessed from left menu via anchor redirects within page
    • Updated synonomies per Trichoptera World Checklist

Page created: November 1, 2012; Last updated: November 05, 2013 (EB)