Aquatic Insects of Michigan

by Ethan Bright, Museum of Zoology Insect Division and School of Natural Resources and Environment
University of Michigan

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Aquatic Ephydridae - Shore and Dance Flies

This dipteran family of about 2,000 species world-wide have largely aquatic immature stages, and with very small to small adults that often not stray far from their aquatic habitats. Aquatic larvae are typically found in shallow areas of muddy margins of freshwater habitats, feeding on small sized algal and plant as well as decaying animal material (Foote, 1995; Courtney & Merritt 2007). Some groups have adapted to saline habitats (salt marshes, tidal pools, salt or alkaline lakes) where they are sometimes constitute the dominant aquatic insect group.

Locality information is based on records at UMMZ-Insect Division and literature citations. Genera are denoted in green; species denoted in bold blue have been recorded in Michigan; other species denoted in black bold are expected to occur based on their known distribution. You can use the Find function in your web browser to locate families, genera, and species. Synonomous species (indented) are listed below the current valid species (denoted in bold) with its taxonomic status indicated.

    Allotrichoma Becker, 1896: 121 (Gymnomyzinae)
    Lentic-temporary puddles (near dung), lotic-margins - burrowers (in dung, temporary puddles, and decaying snails) - collectors-gatherers (dung and decaying snails)
      Allotrichia (Pseudohecamede) abdominale (Williston, 1896) Hecamede (orig.) - OH (Mathis, 1991: 8-14)
      Allotrichoma (Neotrichoma) atrilabre Cresson, 1926: 252 - Mathis and Zatwarnicki 2012: 82-89 (Cheboygan: Bryant's Bog (45°32.8'N, 84°40.3'W), 16 Jun 1933, G. Grant (1m; USNM)
        (syn.) Allotrichoma yosemite Cresson, 1926: 252
      Allotrichoma (Allotrichoma) bezzii Becker, 1896: 123 - west to ne. MN, expect in w. UP/Isle Royale
      Allotrichoma (Allotrichoma) occidentale Mathis & Zatwarnicki, 2012: 50 - Mathis and Zatwarnicki 2012: 50-55 (Iosco Co., 44°20'N, 83°38'W, July 03, 1950, G. C. Steyskal (1?; USNM))
      Allotrichoma (Allotrichoma) simplex (Loew, 1861b: 354 as Discocerina) - Mathis and Zatwarnicki 2012: 65-73 (Monroe Co., 41°55'N, 83°23.9'W, 2 Jul 1939, G. C. Steyskal (1m, 4f; USNM); Newaygo Co., 43°25.2'N, 85°48'W, 27 Jun 1953, R. R. Dreisbach (1?; USNM); Wayne Co., Detroit, 42°23'N, 83°06.1'W, 17 Sep 1939, G. C. Steyskal (1m; USNM); Wayne Co., Grosse Ile, 42°07.7'N, 83°08.7'W, 8 Aug 1954, G. C. Steyskal (2m; USNM))
        (syn.) Allotrichoma filiforme Becker, 1896: 123
        (syn.) Allotrichoma trispinum Becker, 1896: 124
        (syn.) Allotrichoma trispinum of authors, not Becker, 1896
        (syn.) Allotrichoma dahli Beschovski, 1966b: 937

    Athyroglossa Loew, 1860a: 12 (Gymnomyzinae)
    Lentic-littoral, vascular hydrophytes, decaying organic material - burrowers - collectors-gatherers
      Athyroglossa dinorata Mathis & Zatwarnicki, 1990: 892 - Mathis & Zatwarnicki 1990: 894
        (syn.) Athyroglossa ordinata of various American authors, not Becker, 1896: 135 - Wirth, 1965a: 735
      Athyroglossa glabra (Meigen, 1830: 69 as Notiphila) - Wirth 1965: 735
      Athyroglossa granulosa (Cresson, 1922a: 341 as Ochtheroidea) - Wirth 1965: 735; IA, OH, east (Sheiring & Foote, 1973: 158)

    Atissa Haliday, in Curtis 1837: 281 (Hydrellinae)
    Lentic-littoral (saline lakes and pools)
      Atissa humilis (Coquillett, 1902: 183 as Parephydra)
      Atissa litoralis (Cole, 1912: 160 as Allotrichoma) - OH (Sheiring and Foote, 1973: 158)
      Atissa subnuda (Sturtevant & Wheeler, 1954: 252 as Pelignellus)

    Axysta Haliday, 1839: 406 (Ilytheinae)
    Lentic-littoral - sprawlers - collectors-gatherers
      Axysta extera (Cresson, 1924: 162 as Lytogaster) - Krivosheina and Mathis 2010
        (comb.) Microlytogaster extera Clausen, 1983
      Axysta nigrifacies (Miyagi, 1977: 80 as Hyadina) - AK, MN, OH, ON, widespread
        (syn.) Axysta americana Clausen, 1983: 66

    Brachydeutera Loew, 1862c: 162 (Ephydrinae)
    Lentic-littoral - sprawlers, burrowers, swimmers - collectors-gatherers
      Brachydeutera argentata (Walker, 1852: 407 as Notiphila) - Mathis 1983
        (syn.) Brachydeutera neotropica Wirth, 1964: 8 [partial misid.]
        (syn.) Brachydeutera dimidiata Loew, 1862c: 163

    Callinapaea Sturtevant & Wheeler, 1954: 220 (Ephydrinae)
      Callinapaea aldrichi (Sturtevant & Wheeler, 1954: 220 as Napaea) - widespread
      Callinapaea laurentiana (Wirth, 1965b: 63 as Parydra (Callinapaea)) - Clausen & Cook 1971, fig. 234 (depicts record from Isle Royale, but not in text [error?])

    Coenia Robineau-Desvoidy, 1830: 800 (Ephydrinae)
    Lentic, margins - sprawlers, burrowers - collectors-gatherers
      Coenia curvicauda (Meigen, 1830: 116 as Ephydra) - UMMZ-Insect Division; Mathis 1975: 75 (Map 3)

    Diclasiopa Hendel, 1917: 42 (Gymnomyzinae)
    Lentic-littoral, lotic-depositional
      Diclasiopa lacteipennis Loew, 1862: 145 - Mathis and Mathis 2008: 180
        (syn.) Hecamede xanthocera Loew, 1869

    Discocerina Macquart, 1835: 527 (Gymnomyzinae)
    Lentic and lotic margins (primarily terrestrial) - burrowers (in moss and algae) - collectors-gatherers
      Discocerina buccata (Cresson, 1930a: 78 as Hecamedoides (orig.) - Mathis & Zatwarnicki 2013: 153 (Bay: Bay City (43°35.7'N, 83°53.3'W), 30 May 1934, G.C. Steyskal (1m; ANSP). Clare: Clare (43°49.2'N, 84°46.1'W), 12 Jul 1941, R.R. Dreisbach (1?; USNM). Hillsdale: Somerset (42°02.9'N, 84°22.6'W), 12 May 1952, A.H. Sturtevant (1m; USNM). Wayne: Grosse Ile (42°07.7'N, 83°08.7'W), 8 Aug 1954, G.C. Steyskal (5m, 8f; USNM))
        (comb.) Hydrochasma buccatum (Cresson, 1930a) Cresson 1942a: 113
      Discocerina obscurella (Fallén, 1813: 251 as Notiphila) - Mathis & Mathis 2008: 180
        (syn.) Discocerina parva Loew, 1862: 146
        (syn.) Discocerina parva var. nigriventris Cresson, 1916: 148

    Ditrichophora Cresson, 1924: 159 (Gymnomyzinae)
    Lentic-littoral
      Ditrichophora atrata Cresson, 1940: 7 - widespread (Wirth 1965: 739)
      Ditrichophora exigua Cresson, 1924: 159 - Wirth, 1965a: 739
      Ditrichophora simiaceps Cresson, 1940: 6 - IN, west (Wirth 1965: 739)
      Ditrichophora tacoma Cresson, 1924: 160 - widespread (Wirth 1965: 739)

    Ephydra Fallén, 1810: 22 (Ephydrinae)
    Lentic-littoral (in algal mats), lotic-depositional - sprawlers - shredders-herbivores (macroalgae), collectors-gatherers
      Ephydra (Halephydra) cinera Jones 1906: 159 - IL (Steinly et al. 1987: 167), OH (Scheiring and Foote 1973: 163)
        (syn.) Ephydra gracilis Packard, 1871: 105
      Ephydra (Ephydra) millbrae Jones, 1906: 155 - UMMZ-Insect Division [doubtful: probably an error, as Wirth 1971a: 365 and Mathis and Marinoni 2016: 92 note its distribution is western and Australasian/Oceanian]
        (syn.) Ephydra riparia of authors, not Fallén (misid.)
      Ephydra niveiceps Cresson, 1916 - OH (Scheiring and Foote 1973: 163)
      Ephydra (Ephydra) obscuripes Loew, 1866 - UMMZ-Insect Division; distribution widespread (Wirth 1971a: 370)
      Ephydra oscitans Walker, 1849: 1106
        (syn.) Ephydra oscitans Walker, 1858: 253 [preoccupied, Walker, 1849]
      Ephydra (Ephydra) packardi Wirth, 1971a: 365 - IL, IN, widespread (Mathis & Marinoni 2016: 93-94)
      Ephydra (Ephydra) riparia Fallén, 1813: 246 - UMMZ-Insect Division; Wirth 1971: 366
        (syn.) Ephydra halophila Packard, 1869: 49
        (syn.) Ephydra millbrae Jones, 1906: 155
        (syn.) Ephydra lata Walker, 1858: 233

    Glenanthe Haliday, 1839: 404 (Hydrellinae)
      Glenanthe interior Chillcott, 1964: 811 - Mathis, 1995: 8
      Glenanthe salina Mathis, 1995: 15 - OH, widespread (AK-NS-TX) Mathis 1995: 15, Fig. 29

    Gymnoclasiopa Hendel, 1930: 136 (Gymnomyzinae)
      Gymnoclasiopa bohemanni (Becker, 1896: 159 as Clasiopa) - western species, but one record from Ontario north of Lake Superior (Mathis and Zatwarnicki 2012c:25-31)
        (comb.) Discocerina bohemanni Cresson 1925: 256
        (comb.) Ditrichophora bohemanni Hackman 1980: 128
        (comb.) Gymnoclasiopa bohemanni Mathis and Zatwarnicki 1995: 175
        (syn.) Ditrichophora (Gymnoclasiopa) montana Cresson 1942: 120 - Mathis and Zatwarnicki 2012c: 25
        (comb.) Gymnoclasiopa montana Mathis and Zatwarnicki 1995: 177
      Gymnoclasiopa parilis (Cresson, 1924: 160 as Ditrichophora) - widespread north (Mathis and Zatwarnicki 2012c: 42-46)
        (comb.) Gymnoclasiopa parilis Mathis and Zatwarnicki 1995: 177
      Gymnoclasiopa pulchella (Meigen, 1830: 70 as Notiphila) - Mathis and Zatwarnicki 2012c:46-51 (Saginaw: Saginaw (45°25.2'N, 83°57'W), 18 Jun 1952, R. R. Dreisbach (1?; USNM)
        (comb.) Hydrellia pulchella Macquart 1835: 525
        (comb.) Clasiopa pulchella Becker 1905: 195
        (comb.) Discocerina pulchella Czerny 1909: 268
        (comb.) Ditrichophora (Gymnoclasiopa) pulchella Cresson 1942: 121
        (comb.) Gymnoclasiopa pulchella. Mathis and Zatwarnicki 1995: 178
        (syn.) Clasiopa fulgida Becker, 1896: 156
      Gymnoclasiopa tacoma (Cresson, 1924: 160 as Ditrichophora) - Mathis and Zatwarnicki 2012c: 56-61 (Presque Isle: (45.3517°N, -83.4883°W), 2 Jun 1951, R. R. Dreisbach (1?; USNM))

    Haloscatella Mathis, 1979a: 6 (Ephydrinae)
      Haloscatella muria (Mathis, 1979: 12 as Lamproscatella (Haloscatella) - Mathis 1979: 15

    Hecamedoides Hendel, 1917: 41 (Gymnomyzinae)
    Lentic-littoral?
      Hecamedoides unispinosa (Collin, 1943: 148 as Discocerina) - Mathis & Zatwarnicki 2008: 1023-1024 (Fig. 11)
        (comb.) Hecamedoides glaucella var. unispinosa Cogan, 1976: 83
        (mis.id.) Hecamedoides glaucellus of American authors, not Stenhammar, 1844

    Hyadina Haliday, 1839: 406 (Ilytheinae)
    Lotic and lentic - margins - sprawlers - scrapers (graze bluegreen algae)
      Hyadina albovenosa Coquillett, 1900: 34 - Clausen 1983b: 208, 213 (Fig. 31)
      Hyadina binotata (Cresson, 1926: 256 as Hyadina) - Clausen 1983b: 213 (Fig. 32)
        (syn.) Hyadina macquarti Cresson, 1930b: 80
      Hyadina corona (Cresson, 1926: 256 as Hydrina) - Wirth 1965: 751; regional distribution in Clausen 1983b: 214 (Fig. 35)
      Hyadina furva furva (Cresson, 1926: 257 as Lytogaster) - Clausen 1983b: 213 (Fig. 33), 219
      Hyadina subnitida Sturtevant & Wheeler, 1954: 215 - Clausen 1983b: 213 (Fig. 34), 224
      Hyadina vockerothi Clausen, 1984: 83 - ON (Clausen, 1984)

    Hydrellia Robineau-Desvoidy, 1830: 790 (Hydrellinae)
    Lentic-littoral, vascular hydrophytes, lotic-erosional and depositional (vascular hydrophytes-watercress) - burrowers-miners (especially Potamogeton) - shredders-herbivores (miners)
      Hydrellia americana Cresson, 1931: 106 - Deonier 1971: 38
      Hydrellia amnicola Deonier, 1971: 39 - MN (Deonier 1971: 39)
      Hydrellia ascita Cresson, 1942b: 78 - Wirth 1965: 743; Deonier, 1971: 40
      Hydrellia bergi Cresson, 1941: 37 - Cresson, 1941: 37; Deonier, 1971: 42
      Hydrellia bilobifera Cresson, 1936: 262 - Deonier 1971: 44
      Hydrellia borealis Cresson, 1944: 164 - Deonier 1971: 48
      Hydrellia caliginosa Cresson, 1936: 257 - Deonier 1971: 49
      Hydrellia cessator Deonier, 1971: 50 - MB, MN (Deonier 1971: 50)
      Hydrellia columbata Deonier, 1971: 50 - ME, MN (Deonier 1971: 51)
      Hydrellia crassipes Cresson, 1931: 107 - Deonier 1971: 52
      Hydrellia definita Cresson, 1944: 165 - Deonier 1971: 57
      Hydrellia discursa Deonier, 1971: 57 - Deonier 1971: 58
      Hydrellia fascitibia (von Roser, 1940 as Notiphila) - Deonier 1971: 54 (as H. cruralis Coquillett); Mathis and Mathis 2008
        (syn.) Hydrellia cruralis Coquillett, 1910
      Hydrellia formosa Loew, 1861b: 355 - Deonier 1971: 62
      Hydrellia griseola (Fallén, 1813: 250 as Notiphila) - Deonier 1971: 66 (Map 1); Mathis and Mathis 2008
        (comb.) Hydrellia griseola var. hypoleuca Johannsen, 1935: 56
        (comb.) Hydropota griseola Kloet & Hincks, 1945: 396
        (comb.) Hydropota griseola var. scapularis Lange et al., 1953: 8
        (syn.) Hydrellia communis Robineau-Desvoidy, 1830: 790
        (syn.) Hydrellia hypoleuca Lowe, 1862: 151
        (syn.) Hydrellia obscuriceps Lowe, 1862: 152
        (syn.) Hydrellia scapularis Lowe, 1862: 153
      Hydrellia harti Cresson, 1936: 262 - IL, OH, ON s. to Mexico (Deonier 1971: 70; Scheiring and Foote 1973: 163)
      Hydrellia idolator Deonier, 1971: 70 - OH (Scheiring and Foote 1973: 163), ON s. to MS (Deonier 1971: 71)
      Hydrellia ischiaca Loew, 1862: 150 - Deonier 1971: 73
      Hydrellia itascae Deonier, 1971: 76 - MN (Deonier 1971: 76)
      Hydrellia luctuosa Cresson, 1942b: 78 - Deonier 1971: 78
      Hydrellia manitobae Deonier, 1971: 79 - ME-AK (Deonier 1971: 79), expect northern UP
      Hydrellia morrisoni Cresson, 1924: 162 - Deonier 1971: 81
      Hydrellia nobilis (Loew, 1862: 229 as Psilopa) - IL, IN, MN, east (Deonier 1971: 82)
      Hydrellia notiphiloides Cresson, 1924: 162 - Deonier 1971: 85
      Hydrellia penicilli Cresson, 1944: 168 - Deonier 1971: 85
      HydreFllia proctori Cresson, 1934a: 235 - Deonier 1971: 90
      Hydrellia pulla Cresson, 1931: 108 - Deonier 1971: 92
      Hydrellia saltator Deonier, 1971: 94 - ON (Lambton Co.) (Deonier 1971: 94)
      Hydrellia surata Deonier, 1971: 98 - MN, ON, widespread (Deonier 1971: 98)
      Hydrellia suspecta Cresson, 1936: 258 - ME, IA (Deonier 1971: 99)
      Hydrellia tibialis Cresson, 1917: 341 - Deonier 1971: 101
      Hydrellia trichaeta Cresson, 1944: 7 - Wirth, 1965a: 744

    Hydrochasma Hendel, 1936: 101 (Gymnomyzinae)
    Lentic-littoral, lotic-margins
      Hydrochasma avanae Mathis and Zatwarnicki 2010: 113 - Mathis & Zatwarnicki 2013: 117 (Kalamazoo: Kalamazoo (42°17.5'N, 85°35.2'W), 10 May 1936, C. W. Sabrosky (1?; ANSP)
      Hydrochasma leucoproctum (Loew, 1861: 355 as Discocerina) - Mathis et al. 2013: 134, 136 (Bay: Bay City (43°35.7'N, 83°53.3'W), 16 Aug 1936, R. R. Dreisbach (2?, 1?; ANSP). Kalamazoo: Kalamazoo (42°17.5'N, 85°35'W), 10 May 1936, C. W. Sabrosky (1?; ANSP). Monroe: Monroe (41°55'N, 83°23.9'W), 2 Jul 1939, G. C. Steyskal (1?; ANSP).)

    Ilythea Haliday, 1839: 408 (Gymnomyzinae)
      Ilythea spilota (Curtis, 1832: pl. 413 as Ephydra) - UMMZ-Insect Division

    Lemnaphila Cresson, 1933: 229 (Hydrellinae)
    Lentic-vascular hydrophytes (floating zone-uckweed) - burrowers-miners of Lemna (duckweed) - shredders-herbivores
      Lemnaphila scotlandae Cresson, 1933: 229 - Wirth, 1965a; Mathis and Edminston 2000: 667 (Clinton Co.: Bath, 10-Oct-1965 [USNM])

    Leptopsilopa Cresson, 1922: 136 (Discomyzinae)
    Lentic-littoral - burrowers, in decaying vegetation - collectors-gatherers
      Leptopsilopa atrimana (Loew, 1878: 197 as Psilopa) - UMMZ-Insect Division; widespread North America (Wirth 1965: 741)

    Limnellia Malloch, 1925: 331 (Ephydrinae)
    Lentic-littoral
      Limnellia anna Cresson, 1935: 363 - Mathis 1978: 265-266
        (comb.) Scalella (Limnellia) anna Sturtevant & Wheeler, 1954: 177
      Limnellia stenhammari Zetterstedt, 1846: 1842 as Ephydra) - IL, IN, MN, OH, ON, widespread
        (comb.) Scatella stenhammari Loew, 1860: 40
        (comb.) Eustigoptera stenhammari Cresson, 1930a: 128
        (comb.) Stranditella stenahmmari Duda, 1942: 30
        (comb.) Scatella (Limnellia) stenhammeri Sturtevant & Wheeler, 1954: 177
        (syn.) Ephydra oscitans Walker, 1849: 1106
        (syn.) Scatella oscitans Osten Sacken, 1878: 204
        (syn.) Limnellia oscitans Wirth, 1965: 758

    Lytogaster Becker, 1896: 202 (Ilytheinae)
    Lotic and lentic - margins - sprawlers - scrapers (graze bluegreen algae)
      Lytogaster abdominalis (Stenhammar, 1844: 238 as Notiphila sectio Philygria) - UMMZ-Insect Division; Clausen, 1982: 408, 422
      Lytogaster excavata Sturtevant & Wheeler, 1954: 213 as Hyadina gravida excavata) - Clausen, 1982: 413, 422
      Lytogaster perplexa Clausen, 1982: 425 - ON, widespread (Clausen, 1982: 422)

    Mosillus Latreille, 1804: 196 (Gymnomyzinae) Lentic-littoral, (including alkaline lakes and ponds)
      Mosillus bidentatus (Cresson, 1926: 249 as Gymnopa) - Mathis et al. 1993: 10 (Keweenaw Co., Isle Royale, 3-7 Aug 1936, C.W.Sabrosky (USNM); Wayne Co., Detroit, 16 Aug 1930, G.C. Steyskal (ANSP, USNM).
      Mosillus tibialis Cresson, 1916b: 149 - MN, IN, QE-CA (Mathis et al. 1993: 18-22, one record from Indiana Dunes, nw. Indiana; expect in sw. LP)

    Nostima Coquillett, 1900: 35 (Ilytheinae)
    Lotic and lentic - margins - sprawlers - scrapers
      Nostima approximata Sturtevant & Wheeler, 1954: 240 - OH (Edmiston & Mathis 2005: 14)
      Nostima picta (Fallén, 1813) Notiphila (orig.) - Edmiston and Mathis 2005: 58 (Berrien Co., Stevensville, 29 May 1938, C.W. Sabrosky (19; ANSP); Ionia Co., Lyons, 10 Aug 1941, C.W. Sabrosky; Iosco Co., 3 Jul 1950, G.C. Steyskal (2 9; USNM); Menominee Co., Stephenson (19; USNM); Roscommon Co., Roscommon, 20 Jun 1953, R.R. Dreisbach (19; USNM); Wayne Co., Grosse Ile, 5 Oct 1947, GC. Steyskal (19; USNM); Wexford Co., Cadillac, 15 Jun 1941, C.W. Sabrosky (19; USNM))
        (comb.) Ephydra (Hydrina) picta Haliday, 1839
        (comb.) Philygria picta Loew, 1860
        (comb.) Philygria picta var. nigripes Strobl, 1880
        (comb.) Philygriola picta Hendel, 1917
        (comb.) Nostima (Philygriola) picta Cresson, 1930a
        (comb.) Nostima picta picta Mathis & Zatwarnicki, 1995
        (comb.) Nostima picta nigripes Mathis & Zatwarnicki, 1995
        (syn.) Philygria pullula Becker, 1905
        (syn.) Notiphila pullula Fallén, 1823
        (syn.) Nostima (Philygriola) scutellaris Cresson, 1933
        (syn.) Nostima scutellaris scutellaris Sturtevant & Wheeler, 1954
      Nostima quinquenotata Cresson, 1930a: 79 - Edmiston & Mathis 2005: 64-65 (Lapeer Co., Lapeer (21 km N), 30 May 1937, C.W. Sabrosky (19; ANSP).

    Notiphila Fallén, 1810: 22 (Hydrellinae)
    Lentic-littoral (detritus, vascular hydrophytes, bogs), lotic-depositional - burrowers (attached to roots by respiratory spine) - collectors-gatherers and filterers
      Notiphila adusta Mathis, 1979b: 19 - Mathis 1979b: 20
      Notiphila atripes Cresson, 1917b: 50 - UMMZ-Insect Division
      Notiphila avia Loew, 1878: 193 - Mathis 1979b: 30-31
      Notiphila bella Loew, 1862: 135 - Wirth 1965: 747 (as N. nudipes)
        (syn.) Notiphila nudipes Cresson [in part], 1917b: 43
      Notiphila biseriata Cresson, 1917b: 46 - OH (Mathis 1979b: 23)
        (syn.) Notiphila (Notiphila) loewi Cresson, 1946 [in part]
        (syn.) Notiphila (Notiphila) loewi var. biserata Wirth, 1965b [in part]
      Notiphila carinata Loew, 1862: 137 - IN, OH, ON (Mathis and Mathis 2008: 179; Wirth 1965: 746)
      Notiphila caudata Fallén, 1813: 249 - Mathis and Mathis 2008: 179
      Notiphila cressoni Mathis, 1979b: 39 - IL, OH, ON (Mathis 1979b: 40)
      Notiphila footei Mathis, 1979b: 24 - Mathis 1979b: 25
      Notiphila kentensis Huryn, 1987 - OH
      Notiphila loewi Cresson, 1917b: 44 - UMMZ-Insect Division; Mathis 1979b: 45
      Notiphila macrochaeta Loew, 1878: 192 - western to IA, ne. MN, GA (may occur in western UP) (Mathis 1979b: 78)
      Notiphila nudipes Cresson, 1917b: 43 - Mathis 1979b: 26
      Notiphila olivacea Cresson, 1917b: 52 - UMMZ-Insect Division; Mathis 1979b: 82-83
      Notiphila pallidipalpis Cresson, 1940: 8 - Mathis 1979b: 84-85
      Notiphila pauroura Mathis, 1979b: 47 - e. NA (Mathis 1979b: 48)
      Notiphila phaeopsis Mathis, 1979b: 33 - Mathis 1979b: 34
      Notiphila poliosoma Mathis, 1979b: 49 - OH, WI (Mathis 1979b: 49)
      Notiphila pulchra Mathis, 1979b: 34 - Mathis 1979b: 35
        (syn.) Notiphila (Notiphila) riparia North American author [not Meigen, 1830
      Notiphila scalaris Loew, 1862: 134 - UMMZ-Insect Division; Mathis 1979b: 92
      Notiphila shewelli Mathis, 1979b: 50 - Mathis 1979b: 50
      Notiphila sicca Cresson, 1940: 8 - MN, OH (Mathis 1979b: 94)
      Notiphila solita Walker, 1853: 406 - Mathis 1979b: 52
        (comb.) Notiphila (Agrolimna) solita Wirth, 1965a: 748
        (syn.) Notiphila vittata Loew, 1862: 136
      Notiphila taenia Mathis, 1979b: 28 - Mathis 1979b: 28; MSU - Cook Museum (Holotype + Allotype, Mathis 1979)
      Notiphila uliginosa Haliday, 1839: 222 - Mathis 1979b: 96
        (syn.) Notiphila (Agrolimna) olivacea Cresson [in part], 1917: 52

    Ochthera Latreille, 1802: 462 (Gymnomyzinae)
    Lentic-littoral, lotic-depositional and margins - burrowers - predators (engulfers of chironomids)
      Ochthera anatolikos Clausen, 1977: 468 - UMMZ-Insect Division; Clausen, 1977: 472; Mathis and Mathis 2008: 182
      Ochthera borealis Clausen, 1977: 476 - UMMZ-Insect Division; Clausen, 1977: 480
      Ochthera collina Clausen, 1977: 484 - Clausen, 1977: 486
      Ochthera exsculpta Loew, 1862: 160
      Ochthera mantis (De Geer, 1776: 143 as Musca) - UMMZ-Insect Division
      Ochthera tuberculata Loew, 1862: 161 - IL, MA (Wirth 1965: 753), OH (Scheiring and Foote 1973: 163), maybe found in s. LP

    Paracoenia Cresson, 1935: 356 (Ephydrinae)
    Lentic-littoral - sprawlers - scrapers (graze bluegreen algae), collectors-gatherers of fecal material
      Paracoenia (Paracoenia) bisetosa (Coquillett, 1902: 356 as Coenia) - Mathus 1975: 75 (Map 3)
      Paracoenia (Paracoenia) fumosalis Cresson, 1935: 356 - Mathus 1975: 73 (Map 2)

    Paralimna Loew, 1862c: 138 (Hydrellinae)
    Lentic-littoral (margins in sediments near dung) - sprawlers - collectors-gatherers
      Paralimna (Phaiosterna) decipiens Loew, 1878: 195 - Wirth 1965: 748; Ale-Roche & Mathis 2015: 116 (questionable: see range given by Mathis and Zatwarnicki 2002: 72-76, and Ale-Roche & Mathis 2015: 116)
      Paralimna (Paralimna) punctipennis (Wiedemann, 1830: 590 as Notiphila) - UMMZ-Insect Division; Ale-Roche & Mathis 2015: 79 [Allegan: Saugatuck (42°39.3'N, 86°12.1'W), 25 Jun 1938, C. W. Sabrosky (1m, ANSP). Livingston: Hamburg (42°26.9'N, 83°49.1'W), 21 May 1939, G. C. Steyskal (1f; ANSP). Midland: Midland (43°36.8'N, 84°14.9'W), 6-7 Jun 1936, C. W. Sabrosky (1m, 1f; ANSP).]
        (syn.) Paralimna appendiculata Loew 1862c:138

    Parydra Stenhammar, 1844: 187 (Ephydrinae)
    Lentic-littoral, lotic-depositional and margins - burrowerers - scrapers

      Parydra (Chaetoapnaea) alpina (Cresson, 1924: 163 as Napaea) - Wirth 1965; Clausen & Cook 1971: 35
      Parydra (Chaetoapnaea) appendiculata Loew, 1878: 202 - UMMZ-Insect Division; Clausen and Cook 1971: 69, fig. 229
      Parydra (Parydra) aquila aquila (Fallén, 1813: 247 as Ephydra) - UMMZ-Insect Division; Clausen and Cook 1971: fig. 223
        (syn.) Parydra bituberculata Loew, 1862: 165
        (syn.) Napaea (Parydra) bituberculata bituberculata (Loew), Sturtevant & Wheeler 1954: 227
        (syn.) Parydra nitida Cresson, 1915: 70
        (syn.) Napaea (Parydra) bituberculata nitida (Cresson), Sturtevant & Wheeler 1954: 227
      Parydra (Chaetopanaea) borealis (Cresson, 1949: 235) - UMMZ-Insect Division; Clausen & Cook 1971: 74 (specimen from Isle Royale, C. Sabrosky, fig. 234?)
      Parydra breviceps Loew, 1862: 167 - UMMZ-Insect Division; Clausen & Cook 1971: 45, fig. 224
        (syn.) Parydra limpidipennis Loew, 1878: 201
        (syn.) Parydra vicina Cresson, 1940: 9
        (syn.) Napaea (Parydra) breviceps breviceps (Loew), Sturtevant & Wheeler 1954: 223 (syn.) Napaea (Parydra) breviceps vicina (Cresson), Sturtevant & Wheeler 1954: 223
      Parydra (Parydra) fossarum Haliday, 1833: 175 as Ephydra) - Mathis and Zatwarnicki 1990: 901; Clausen and Cook 1971: 85, fig. 230
        (syn.) Parydra (Chaetoapnaea) parasocia Clausen in Clausen and Cook, 1971: 83
      Parydra (Chaetoapnaea) paullula Loew, 1862: 167 - UMMZ-Insect Division; Clausen and Cook 1971: 86, fig. 233
      Parydra (Chaetoapnaea) quadriloba Clausen, in Clausen & Cook, 1971: 90 - Clausen and Cook, 1971: 92, fig. 225
      Parydra (Parydra) quadrituberculata Loew, 1862: 165 - UMMZ-Insect Division; Clausen and Cook 1971: fig. 226
        (syn.) Napaea (Parydra) quadrituberculata (Loew), Sturtevant & Wheeler 1954: 228
      Parydra (Chaetopanaea) vulgaris (Cresson, 1949: 233) - widespread boreal (Clausen & Cook 1971), maybe be found in boreal habitats of UP or Isle Royale)

    Pelina Haliday, 1839: 407 (Ilytheinae)
    Lotic and lentic - margins - sprawlers - scrapers
      Pelina canadensis Cresson, 1934b: 208 - northwest to MN, OH, WI (Clausen 1973: 131)
      Pelina compar Cresson, 1934b: 207 - Clausen 1973: 132, 154 (Fig. 74) (Cresson 1940, holotype male of P. rudis from Breveort, 02-Aug-1936, C. Sabrosky)
        (syn.) Pelina rudis Cresson, 1940: 9
      Pelina neaenescens Clausen, 1973: 133 - Clausen 1973: 133, paratype from Benzie Co.: Honor, Mich, 9-Jul-1941, C.W. Sabrosky [USNM])
      Pelina truncatula truncatula Loew, 1878: 198 - UMMZ-Insect Division; Clausen 1973: 139, 149, 154 (Fig. 75)

    Philygria Stenhammar, 1844: 154 (Ilytheinae)
    Lotic and lentic - margins - sprawlers - scrapers
      Philygria debilis Loew, 1861: 357 - UMMZ-Insect Division
      Philygria punctatonervosa (Fallén, 1813: 254 as Notiphila) - Mathis and Mathis 2008
        (comb.) Hydrina punctatonervosa Becker, 1926: 57
        (syn.) Philygria opposita Loew, 1861: 356

    Platygymnopa Wirth, 1971b: 266 (Gymnomyzinae)
    Lotic and lentic margins - burrowers (in decomposing snails) - collectors-gatherers (decomposing snails)
      Platygymnopa helicis Wirth, 1971b: 266 - widespread northern

    Polytrichophora Cresson, 1924: 161 (Gymnomyzinae)
    Lentic-littoral, lotic-margins
      Polytrichophora conciliata Cresson, 1924: 161 - Mathis and Zatwarnicki 2012a
      Polytrichophora orbitalis (Loew, 1861: 354 as Discocerina) - Mathis and Zatwarnicki 2012a: 91: 54
      Polytrichophora sturtevantorum Mathis & Zatwarnicki, 2012a: 101 - IL, OH, WI (Mathis and Zatwarnicki 2012a)

    Pseudohyadina Clausen, 1983b: 224 (Ilytheinae)
      Pseudohyadina longicornis Sturtevant & Wheeler, 1954: 214 as Hyadina) - OH, ON (maybe found in se. LP)

    Psilopa Fallén, 1823: 6 (Discomyzinae)
    Lentic-littoral, vascular hydrophytes - planktonic, burrowers - collectors-gatherers, shredders, miners
      Psilopa compta (Meigen, 1830: 68 as Notiphila) - AK-QB,k s. to CA-VA
        (nom.nud.) Psilopa gaudens Cresson in Strickland, 1946: 167
      Psilopa dupla Cresson, 1940: 2 - IL (Steinly et al. 1987: 167), OH (Scheiring and Foote 1973: 163), IA, east and south
      Psilopa girschneri Von Röder, 1889: 54 - widespread
        (syn.) Psilopa olga Cresson, 1922: 137
      Psilopa leucostoma (Meigen, 1830: 68 as Notiphila) - widespread

    Ptilomyia Coquillett, 1900: 261 (Hydrellinae)
    lentic-littoral
      Ptilomyia parva (Williston, 1896: 399 as Hydrellia) - OH (Sheiring and Foote 1973: 158 as P. enigma, Mathus 1997)
        (syn.) Ptilomyia enigma Coquillett, 1900: 262

    Rhysophora Cresson, 1924: 159 (Discomyzinae)
    Lentic-littoral, vascular hydrophytes - burrowers? - shredders-herbivores (miners?)
      Rhysophora robusta Cresson, 1924: 159 - Wirth, 1965a: 743
        (nom.nud.) Discocerina magna Coquillett in Johnson, 1910: 806

    Scatella Robineau-Desvoidy, 1830: 801 (Ephydrinae)
    Lentic-vascular hydrophytes and algal mats, lotic-margins - sprawlers-climbers - collectors-gatherers, scrapers (graze algae)
      Scatella crassicosta Becker, 1896: 234 - northern
      Scatella favillacea Loew, 1862: 170 - widespread
      Scatella laxa Cresson, 1933: 70 - widespread
      Scatella obscuriceps Cresson, 1915: 72
      Scatella obsoleta Loew, 1861b: 358 - IL (Steinly et al. 1987: 167), widespread
      Scatella paludum (Meigen, 1830: 118 as Ephydra) - ON, widespread
      Scatella picea (Walker, 1849: 1105 as Ephydra) - UMMZ-Insect Division
      Scatella quadrinotata Cresson, 1933: 70 - IL (Steinly et al. 1987: 167), widespread
      Scatella setosa Coquillett, 1900: 462
        (syn.) Scatella intermedia Cresson, 1915: 72
      Scatella stagnalis (Fallén, 1813: 248 as Ephydra) - UMMZ-Insect Division

    Scatophila Becker, 1896: 237 (Ephydrinae)
    Lentic-littoral, lotic-margins - sprawlers - scrapers
      Scatophila arenaria Cresson, 1935: 372 - MN, west
      Scatophila despecta (Haliday, 1839: 409 as Ephydra) - ME, ON, widespread north
        (syn.) Scatophila striata Walker, 1849: 1107
        (syn.) Scatophila variabilis Cresson, 1917: 341
      Scatophila mesogramma (Loew, 1869: 42 as Scatella) - RI-AK
      Scatophila unicornis Czerny, 1900: 205 - IL (Steinly et al. 1987: 167), ON, non-native introduced from Europe
        (syn.) Scatophila tuberculosa Cresson, 1935a: 368

    Setacera Cresson, 1930a: 116 (Ephydrinae)
    Lentic-littoral (algal mats), lotic-margins sprawlers - shredders-herbivores, scrapers (graze algae)
      Setacera atrovirens Loew, 1862c: 169 as Ephydra) - Mathis 1982: 13
      Setacera pacifica (Cresson, 1925: 167 as Ephydra (orig.) - Mathis 1982: 48

    Trimerina Macquart, 1835: 528 (Discomyzinae)
    Lentic, vascular hydrophytes (emergent zones) - climbers - predators (piercers, spider egg masses [Foote 1984])
      Trimerina madizans (Fallén, 1813: 253 as Notiphila) - UMMZ-Insect Division; OH (Foote 1984: 486)

    Typopsilopa Cresson, 1916: 147 (Hydrellinae)
    Lentic-littoral (detritus at margins in emergent vascular hydrophyte beds) - sprawlers - collectors-gatherers
      Typopsilopa atra (Loew, 1862: 143 as Psilopa) - Wirth, 1968: 231
        (syn.) Psilopa scoriacea Loew, 1962: 142 - Wirth 1965: 743

    Zeros Cresson, 1943: 10 (Ilytheinae)
    Lotic-depositional (detrital sediments) - sprawlers - collectors-gatherers
      Zeros flavipes (Williston, 1896: 403 as ?Ilythea) - IA, OH (Scheiring and Foote, 1973: 160), ON, southward to Neotropical (Wirth, 1965a: 746)

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Page Updates:

  • May 13, 2017
    • Major corrections and additions to nomenclature, organization and citations, removal of taxa not occurring in the region.
  • December 30, 2013
    • Incorporated information from Mathis & Zatwarnicki 2012 on Gymnoclasiopa

Page created: April 25, 2006 - Last updated: May 13, 2017 (EB)